Eryx Colubrinus
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''Eryx colubrinus'', also known as the Kenyan sand boa Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . and several other
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
s, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Erycinae The Erycinae, also known as the Old World sand boas, are a subfamily of nonvenomous snakes in the family Boidae. Species of the subfamily Erycinae are found in Europe, Asia Minor, Africa, Arabia, central and southwestern Asia, India, Sri Lanka, ...
of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Boidae The Boidae, commonly known as boas or boids, are a family of nonvenomous snakes primarily found in the Americas, as well as Africa, Europe, Asia, and some Pacific islands. Boas include some of the world's largest snakes, with the green anaconda ...
. The species is native to northern and eastern Africa. Three
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognized.


Description

''Eryx colubrinus'' is a heavily-built snake with a small head, small eyes, vertical pupils, and a short tail. Scale texture is extremely smooth, except on the tail, which is covered in bumps. Adult female specimens of ''G. colubrinus'' are rarely more than 91 cm (3 feet) in total length (tail included). The average Egyptian sand boa grows no longer than 12-24" (30-60cm) long, with males being significantly smaller than females. The color pattern usually consists of a yellow or orange coloration overlaid with dark brown splotches. The belly is white or cream-colored. ''E. colubrinus'' is readily available in the pet trade due to its small size, docility and ease of care. In recent years, there have been a number of new
color morph Color (or colour in Commonwealth English; see spelling differences) is the visual perception based on the electromagnetic spectrum. Though color is not an inherent property of matter, color perception is related to an object's light absor ...
s made available by both commercial and hobby breeders. Some of the more popular morphs available include anerythristic (black and white lacking orange/red simple
recessive In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and ...
trait), albino (lacking black pigment simple recessive trait), snow (double recessive combination of an anerythristic and an albino), stripes (normal-colored, anerythristic, albino and snow), hypo/ghost, paradox albinos (simple recessive trait), paradox snows (double recessive trait), splash (recessive trait), paint (recessive trait) and stripe combinations with any of the listed recessive traits. In addition, many line bred traits have been accentuated on the above morphs, such as Nuclears (extreme red), High Whites and Reduced Patterns, for example. File:Gongylophis colubrinus loveridgei.jpg, ''loveridgei'' subspecies "normal" morph File:Eryx colubrinus albino morph.jpg, Albino morph File:Stripe morph Eryx colubrinus.jpg, Stripe morph File:Eryx colubrinus stripe morph lateral view.jpg, Lateral view of a stripe morph showing the specialized head shape for burrowing


Common names

Common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
s for ''Eryx colubrinus'' include East African sand boa, Egyptian sand boa, Kenya sand boa, Kenyan sand boa, and sand boa.


Geographic range

''Eryx colubrinus'' is found in
North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...
from
Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...
as far west as
Niger Niger, officially the Republic of the Niger, is a landlocked country in West Africa. It is a unitary state Geography of Niger#Political geography, bordered by Libya to the Libya–Niger border, north-east, Chad to the Chad–Niger border, east ...
(Aïr), including
Somalia Somalia, officially the Federal Republic of Somalia, is the easternmost country in continental Africa. The country is located in the Horn of Africa and is bordered by Ethiopia to the west, Djibouti to the northwest, Kenya to the southwest, th ...
,
Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
,
Sudan Sudan, officially the Republic of the Sudan, is a country in Northeast Africa. It borders the Central African Republic to the southwest, Chad to the west, Libya to the northwest, Egypt to the north, the Red Sea to the east, Eritrea and Ethiopi ...
,
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
, and northern
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
. A single specimen has been reported from
Yemen Yemen, officially the Republic of Yemen, is a country in West Asia. Located in South Arabia, southern Arabia, it borders Saudi Arabia to Saudi Arabia–Yemen border, the north, Oman to Oman–Yemen border, the northeast, the south-eastern part ...
. The type locality given is "Ægypto".


Habitat

''Eryx colubrinus'' occurs in
semi-desert A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of sem ...
and scrub
savannahs A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the Canopy (forest), canopy does not close. The open canopy allows sufficient li ...
and rock outcroppings. It prefers sandy,
friable In materials science, friability ( ), the condition of being friable, describes the tendency of a solid substance to break into smaller pieces under stress or contact, especially by rubbing. The opposite of friable is indurate. Substances tha ...
soil.


Behavior

''Eryx colubrinus'' is most active at night, but may be irregularly active during the day. This species is
fossorial A fossorial animal () is one that is adapted to digging and which lives primarily (but not solely) underground. Examples of fossorial vertebrates are Mole (animal), moles, badgers, naked mole-rats, meerkats, armadillos, wombats, and mole salamand ...
and spends most of its time underground. During the hotter times of the year, ''E. colubrinus'' seeks refuge beneath stones and in the burrows of small mammals. However, they have also been known to occasionally climb trees.


Diet

''Eryx colubrinus'' feeds on small mammals (such as
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s), lizards and birds that are quickly seized when passing within striking range and killed by constriction. Occasionally, they have been known to hunt out the nests of small mammals and birds.


Reproduction

''Eryx colubrinus'' is
ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a "bridging" form of reproduction between egg-laying oviparity, oviparous and live-bearing viviparity, viviparous reproduction. Ovoviviparous animals possess embryos that develo ...
. In captivity, it breeds readily November through April in the United States, delivering live birth averaging 10-20 babies born spring through late summer. The young at birth typically are 20–25 cm (8-10 inches) in length.


Taxonomy

A synonym for this species, ''Eryx colubrinus'', is ''Anguis colubrina'', given by
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
. Stimson (1969) recognized two subspecies: ''Eryx colubrinus colubrinus'' and ''E. c. loveridgei'' , but mentioned that a number of other authors, including Ahl (1933), Loveridge (1936), Scortecci (1939), and
Parker Parker may refer to: People * Parker (given name), including a list of people and fictional characters with the given name * Parker (surname), including a list of people and fictional characters with the surname Arts and entertainment * ''Parke ...
(1949), questioned whether ''E. c. loveridgei'' was valid and considered the species to be monotypic with geographic variation. When recognized, ''Eryx colubrinus loveridgei'' is said to occur in the southern part of the range and is described as being more orange in color. A third subspecies, ''Eryx colubrinus rufescens'', is smaller and more rounded than ''E. c. loveridgei'', with rougher scales. Shi et al. (2023) proposed that ''E. colubrinus'' was the sister species to the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
fossil species '' E. linxiaensis'' from
Gansu Gansu is a provinces of China, province in Northwestern China. Its capital and largest city is Lanzhou, in the southeastern part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibetan Plateau, Ti ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
. This suggests repeat dispersal events of the genus from Africa to Eurasia during the Miocene.


Etymology

The
subspecific name In biology, trinomial nomenclature is the system of names for taxa below the rank of species. These names have three parts. The usage is different in zoology and botany. In zoology In zoological nomenclature, a trinomen (), trinominal name, or ...
''loveridgei'' is in honor of British
herpetologist Herpetology (from Ancient Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is a branch of zoology concerned with the study of amphibians (including frogs, salamanders, and caecilians (Gymnophiona)) and reptiles (in ...
Arthur Loveridge Arthur Loveridge (28 May 1891 – 16 February 1980) was a British people, British biologist and Herpetology, herpetologist who wrote about animals of East Africa, particularly Tanzania, and of New Guinea. He gave Binomial nomenclature, scientific ...
. Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins University Press. xiii + 296 pp. . (''Eryx colubrinus loveridgei'', p. 161).


References


Further reading

* 624 pp. (''Eryx colubrinus'', pp. 376–377).


External links

* {{Taxonbar, from=Q253890 colubrinus Reptiles described in 1758 Taxa named by Carl Linnaeus