Erythrolamprus Aesculapii
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''Erythrolamprus aesculapii'', also known commonly as the Aesculapian false coral snake, the South American false coral snake, and in Portuguese as ''bacorá'', or ''falsa-coral'', is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of mildly venomous snake in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Colubridae Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. C ...
. The species is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
. Freiberg M (1982). ''Snakes of South America''. Hong Kong: T.F.H. Publications. 189 pp. . (''Erythrolamprus aesculapii'', pp. 66, 98, 132 + photograph on p. 42 + line drawing of head scalation on p. 67).


Etymology

The specific name, ''aesculapii'', refers to Aesculapius, the Greek mythological god of medicine, who is depicted with a snake-entwined staff.


Classification

''Erythrolamprus aesculapii'' belongs to the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Erythrolamprus ''Erythrolamprus'' is a genus of colubrid snakes native to Central America, the Caribbean, and South America. They include the false coral snakes, which appear to be coral snake mimics. Classification The genus ''Erythrolamprus'' belongs to ...
'', which contains over 50 species. The genus ''Erythrolamprus'' belongs to the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Dipsadinae Dipsadinae is a large subfamily of colubroid snakes, sometimes referred to as a family (Dipsadidae). Species of the subfamily Dipsadinae are found in most of the Americas, including the West Indies, and are most diverse in South America. There a ...
, which is sometimes referred to as the family Dipsadidae. Molecular
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
DNA analysis has shown that ''Erythrolamprus aesculapii'' is likely
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
, as shown in the
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below of ''Erythrolamprus'' species located in northern South America:


Geographic range

''E. aesculapii'' is found in the
Amazon rainforest The Amazon rainforest, also called the Amazon jungle or Amazonia, is a Tropical and subtropical moist broadleaf forests, moist broadleaf tropical rainforest in the Amazon biome that covers most of the Amazon basin of South America. This basin ...
of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
. It is also found on the island of
Trinidad Trinidad is the larger, more populous island of the Republic of Trinidad and Tobago, the country. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is the southernmost island in ...
(in the Republic of Trinidad and Tobago).


Habitat and behavior

''E. aesculapii'' is often found in the
leaf litter Plant litter (also leaf litter, tree litter, soil litter, litterfall, or duff) is dead plant material (such as leaves, bark, needles, twigs, and cladodes) that has fallen to the ground. This detritus or dead organic material and its constituen ...
or burrowing in the
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
in rain forests, at altitudes from sea level to .


Diet

''E. aesculapii'' feeds mainly on other snakes, including venomous species. It will also prey on lizards, fish and earthworms. Insects are probably consumed though secondary ingestion.


Venom

''E. aesculapii'' is mildly venomous.


Mimicry

The brightly colored, ringed patterns of snakes of the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Erythrolamprus ''Erythrolamprus'' is a genus of colubrid snakes native to Central America, the Caribbean, and South America. They include the false coral snakes, which appear to be coral snake mimics. Classification The genus ''Erythrolamprus'' belongs to ...
'' resemble those of
sympatric In biology, two closely related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter each other. An initially interbreeding population that splits into two or more distinct spe ...
coral snakes of the genus ''
Micrurus ''Micrurus'' is a genus of venomous coral snakes of the family Elapidae. Geographic range Species in the genus ''Micrurus'' are endemic to the Americas. Species The following 82 species are recognized as being valid. *'' Micrurus albicinctus' ...
'', and it has been suggested that this is due to
mimicry In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. In the simples ...
. Whether this is classical
Batesian mimicry Batesian mimicry is a form of mimicry where a harmless species has evolved to imitate the warning signals of a harmful species directed at a predator of them both. It is named after the English naturalist Henry Walter Bates, who worked on butt ...
, classical
Müllerian mimicry Müllerian mimicry is a natural phenomenon in which two or more well-defended species, often foul-tasting and sharing common predators, have come to mimicry, mimic each other's honest signal, honest aposematism, warning signals, to their mutuali ...
, a modified form of Müllerian mimicry, or no mimicry at all, remains to be proven. Goin CJ, Goin OB, Zug GR (1978). ''Introduction to Herpetology, Third Edition''. San Francisco: W.H. Freeman. xi + 378 pp. . (Mimicry in ''Erythrolamprus'', p. 159).


Subspecies

The following four
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognized as being valid, including the
nominotypical subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
: *''Erythrolamprus aesculapii aesculapii'' (
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758)
Amazon River Basin *''Erythrolamprus aesculapii monozonus'' Jan, 1863
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
(
Bahia Bahia () is one of the 26 Federative units of Brazil, states of Brazil, located in the Northeast Region, Brazil, Northeast Region of the country. It is the fourth-largest Brazilian state by population (after São Paulo (state), São Paulo, Mina ...
state to
Rio de Janeiro Rio de Janeiro, or simply Rio, is the capital of the Rio de Janeiro (state), state of Rio de Janeiro. It is the List of cities in Brazil by population, second-most-populous city in Brazil (after São Paulo) and the Largest cities in the America ...
state) *''Erythrolamprus aesculapii tetrazonus'' Jan, 1863 – southwestern
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
*''Erythrolamprus aesculapii venustissimus'' ( Wied, 1821) – eastern Bolivia to southeastern Brazil and northeastern
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...


References


Further reading

* * Linnaeus L (1758). ''Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio Decima, Reformata.'' Stockholm: L. Salvius. 824 pp. (''Coluber æsculapii'', new species, p. 220). (in Latin). * {{Taxonbar, from=Q2710586 aesculapii Fauna of the Amazon Snakes of Brazil Reptiles of Bolivia Reptiles of Argentina Reptiles of Colombia Reptiles of Ecuador Reptiles of French Guiana Reptiles of Peru Reptiles of Paraguay Reptiles of Trinidad and Tobago Reptiles of Venezuela Reptiles described in 1758 Taxa named by Carl Linnaeus