Erynnis Tages
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The dingy skipper (''Erynnis tages'') is a species of
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
in the family
Hesperiidae Skippers are a group of butterflies placed in the family Hesperiidae within the order Lepidoptera (moths and butterflies). They were previously placed in a separate superfamily, Hesperioidea, but have since been placed in the superfamily Papilion ...
.


Description

''Erynnis tages'' is different from other skippers because of the predominantly monochrome, gray-brown wing coloration and the marbling, which is only present on the upper side of the forewings, as well as a series of small white dots on the wing edge. It is probably the most moth-like British butterfly and normally rests with its wings in a moth-like fashion. This well-camouflaged, brown and grey butterfly can be confused with the
grizzled skipper ''Pyrgus'' is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genera '' Burnsius'' and '' C ...
, the Mother Shipton moth or the burnet companion moth. Faded specimens of '' Carcharodus alceae'' are distinguished by glass spots in the forewing and by a jagged rear wing edge. A special feature is the restraint of the imagos, which is reminiscent of a deltoid moth with its roof-shaped wings placed one on top of the other.


Description in Seitz

Th. tages L. (86 c). Forewing grey-brown, with two oblique black bands, which are proximally edged with yellowish grey. A marginal row of small light dots and two apical dots, of which one is transparent. Hindwing black, in quite normal specimens without spots. Throughout Europe and Northern Asia to the Amur. ab. ''clarus'' aradja, 1895 ab.are very pale specimens, which may occur everywhere among true ''tages''. — ''popoviana'' Nordm. (= ''sinina'' Gr.-Grsh.) ow species ''Erynnis popoviana''">Erynnis_popoviana.html" ;"title="ow species ''Erynnis popoviana">ow species ''Erynnis popoviana''(86 c) is hardly more than a synonym; light grey, with a row of white marginal dots and second similar row in the centre, the hindwing with a light discocellular spot. Dauria. Amur, China. — ''unicolor'' Frr. [ now subspecies] (86 c) is uniformly brown grey without any markings, Greece and Asia Minor. — ''cervantes'' Grasl.[now ''E. tages'' ssp. ''cervantes'' Graslin, 1836 ] (86 d). Larger and much darker than true ''tages''. The dark bands obsolete or only indicated by black streaks: the marginal dots scarcely visible. South Spain. — Larva green with the head brown and yellow lateral stripe dotted with black: on ''Eryngium'' and ''Lotus'', in July and late in the autumn. Pupa green, with reddish abdomen. The butterflies in April and May and again from July onward, everywhere plentiful. They fly low above the ground and like to settle on roads.


Taxonomy

Subspecies are little defined and include ''E. t. unicolor'' Freyer, 1852 found in
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and West Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Armenia, ...
and ''E. t. baynesi'' found in
the Burren The Burren ( ; ) is a karst/glaciokarst landscape centred in County Clare, on the west coast of Republic of Ireland, Ireland.
,
County Clare County Clare () is a Counties of Ireland, county in the Provinces of Ireland, province of Munster in the Southern Region, Ireland, Southern part of Republic of Ireland, Ireland, bordered on the west by the Atlantic Ocean. Clare County Council ...
,
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
. ''Synonyms: Erynnis morio'' SCOPOLI, 1763; ''Erynnis geryon'' ROTTEMBURG, 1775 Dingy skipper (Erynnis tages baynesi) male Burren.jpg, Male ''E. t. baynesi''
County Clare, Ireland Dingy skipper (Erynnis tages baynesi) female Burren.jpg, Female ''E. t. baynesi''
County Clare, Ireland The larvæ of the British butterflies and moths BHL41107797.jpg, Figs 3 larva after 2nd moult 3a larva after 3rd moult 3b, 3c larva after 4th moult


Range

The dingy skipper is widespread from the
Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...
and Ireland in the west to the Pacific in East Asia. In the north, the area boundary follows roughly the 62°N. In Asia, the species penetrates south across the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
to Pakistan, and in China the area boundary is far to the south. In
Bavaria Bavaria, officially the Free State of Bavaria, is a States of Germany, state in the southeast of Germany. With an area of , it is the list of German states by area, largest German state by land area, comprising approximately 1/5 of the total l ...
, the distribution shows a noticeable change from more or less densely populated areas to regions in which the species is absent over long stretches. In southern Bavaria, the species is concentrated in the Alpine region, the valleys of the Alpine rivers, in particular the Lech and
Isar The Isar () is a river in Austria and in Bavaria, Germany. Its source is in the Karwendel mountain range of the Alps. The Isar river enters Germany near Mittenwald and flows through Krün, Wallgau, Bad Tölz, Munich, and Landshut before reaching ...
, as well as the areas near the river in the Donaumoos. In Northern Bavaria, the focus is on the
Franconian Alb The Franconian Jura ( , , or ) is an upland in Franconia, Bavaria, Germany. Located between two rivers, the Danube in the south and the Main in the north, its peaks reach elevations of up to and it has an area of some 7053.8 km2. Emil Mey ...
, in parts of the Franconian Keuper-Lias-Land and the Mainfränkische Platten. A secondary focus in the Upper Main hill country leads to scattered evidence along the northeastern border. Here the species occurs from the
Vogtland Vogtland (; ) is a region spanning the German states of Bavaria, Saxony and Thuringia and north-western Bohemia in the Czech Republic. It overlaps with and is largely contained within Euroregio Egrensis. The name alludes to the former leadershi ...
over the
Fichtelgebirge The Fichtel Mountains (, ; ) is a mountain range in Germany and the Czech Republic. They extend from the valley of the Red Main River in northeastern Bavaria to the Karlovy Vary Region in western Czech Republic. The Fichtel Mountains contain a ...
to the
Upper Palatinate The Upper Palatinate (; , , ) is an administrative district in the east of Bavaria, Germany. It consists of seven districts and 226 municipalities, including three cities. Geography The Upper Palatinate is a landscape with low mountains and nume ...
and
Bavarian Forest image:Zell-bayerischer-wald.jpg, The village of Zell in the Bavarian Forest The Bavarian Forest ( or ''Bayerwald'' ; ) is a wooded, low-mountain region in Bavaria, Germany, that is about 100 kilometres long. It runs along the Czech Republic, C ...
. With the exception of the East Bavarian low mountain range, the occurrence of the dingy skipper coincides with the distribution of limestone areas (Jura and Muschelkalk) or geological features (basalt, gypsum, basic sandstones such as dolomitic arkose). It is widely but patchily distributed across Britain. It occurs further north than any other skipper in
Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
with some isolated colonies in the
Inverness Inverness (; ; from the , meaning "Mouth of the River Ness") is a city in the Scottish Highlands, having been granted city status in 2000. It is the administrative centre for The Highland Council and is regarded as the capital of the Highland ...
region. It is also one of the two skippers to be found in Ireland, again with a patchy distribution but the main strongholds along the western side. It is on the decline in several European countries including the UK and
Armenia Armenia, officially the Republic of Armenia, is a landlocked country in the Armenian Highlands of West Asia. It is a part of the Caucasus region and is bordered by Turkey to the west, Georgia (country), Georgia to the north and Azerbaijan to ...
.Butterfly Conservation Armenia
/ref>


Habitat

''Erynnis tages'' favours open grassy habitats up to 2,000 metres above sea level. A variety of habitats are used including
chalk downland Downland, chalkland, chalk downs or just downs are areas of open chalk hills, such as the North Downs. This term is used to describe the characteristic landscape in southern England where chalk is exposed at the surface. The name "downs" is deriv ...
, woodland clearings, coastal dunes, railway lines and waste ground. The habitats of the dingy skipper are mainly dry and poor grasslands. Extensive grassland with one or two-tier meadows and pastures as well as habitats with little vegetation with raw soil and initial plant communities are also regularly populated. In forests, the imagos fly in very sparse wooded stands or on forest meadows, on the edges of paths and forest edges. The species is also found in fens. Suitable habitats are disturbances such as paths with a dry, warm microclimate.


Ecology

The imagos fly in two generations from May-June and July-August but in northern regions and at the high altitudes, there is only a single generation. The flight period in Bavaria extends from the middle of April to the beginning of September with a focus from the beginning of May to the end of June and a maximum in the last May decade. First and second generation overlap. The second generation, which occurs only irregularly or regionally, is always significantly weaker. The imagos prefer low-growing or very patchy vegetation and often stay on bare ground to sunbathe or to absorb moisture and minerals. On the ground or on low plants, the males also move from perch in order to track down the females by approaching insects of suitable size. The flight is usually very fast and low, whereby the locally loyal imagos usually only cover short distances of a few meters and soon settle again. There is very little information on flower visits. In Bavaria out of a total of 14 listed plant species, only three are mentioned several times, in addition to the most important egg-laying plants ''
Hippocrepis comosa ''Hippocrepis comosa'', the horseshoe vetch, is a species of perennial flowering plant belonging to the genus '' Hippocrepis'' in the family Fabaceae. Description The overall appearance depends on its habitat: sometimes it forms upright clump ...
'' and ''
Lotus corniculatus ''Lotus corniculatus'' is a flowering plant in the pea family Fabaceae. Common names include common bird's-foot trefoil, eggs and bacon, birdsfoot deervetch, and just bird's-foot trefoil (a name also often applied to other ''Lotus'' spp.). It ha ...
'', ''
Ajuga reptans ''Ajuga reptans'' is commonly known as bugle, blue bugle, bugleherb, bugleweed, carpetweed, carpet bugleweed, carpet bugle, and common bugle, and traditionally, (however less commonly) as St. Lawrence plant. It is an herbaceous plant, herbaceous f ...
'' also is visited. Other suction plants are yellow-flowered
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
such as ''
Anthyllis vulneraria ''Anthyllis vulneraria'', the common kidneyvetch, kidney vetch or woundwort is a medicinal plant native to Europe, northern Africa, and Western Asia. The name ''vulneraria'' means "wound healer". Description ''Anthyllis vulneraria'' reaches in ...
, Chamaecytisus ratisbonensis'' and '' Medicago falcata'' as well as blue- and violet-flowered mint flowers such as ''
Salvia pratensis ''Salvia pratensis'', the meadow clary or meadow sage, is a species of flowering plant in the family Lamiaceae, native plant, native to Europe, western Asia and northern Africa. The Latin Binomial nomenclature, specific epithet ''pratensis'' ...
'' and ''
Thymus The thymus (: thymuses or thymi) is a specialized primary lymphoid organ of the immune system. Within the thymus, T cells mature. T cells are critical to the adaptive immune system, where the body adapts to specific foreign invaders. The thymus ...
spp.'' Visits to flowers on taller perennials such as ''
Cirsium arvense ''Cirsium arvense'' is a perennial species of flowering plant in the family Asteraceae, native throughout Europe and western Asia, northern Africa and widely introduced elsewhere.Joint Nature Conservation Committee''Cirsium arvense'' The standa ...
'', ''
Eupatorium cannabinum ''Eupatorium cannabinum'', commonly known as hemp-agrimony, or holy rope, is a herbaceous plant in the family Asteraceae. It is a robust Perennial plant, perennial native to Europe, NW. Africa, Turkey, Syria, Iran, Iraq, Jordan, the Caucasus and ...
'' or ''
Echium vulgare ''Echium vulgare'', known as viper's bugloss and blueweed,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Royal Ontario Museum, p. 203. is a species of flowering plant in the borag ...
'' are rare, as the imagos mainly stay near the ground. In addition to visiting flowers, the imagos regularly suckle on moist soil, carrion and excrement. Eggs are laid individually on the upper side of mostly terminal leaflets near the ground, with gaps or weak-growing, microclimatically favored locations being preferred. Observations of egg laying or of egg and caterpillar finds are only available in isolated cases. The following egg-laying and host plants have been reported from Bavaria: ''
Hippocrepis comosa ''Hippocrepis comosa'', the horseshoe vetch, is a species of perennial flowering plant belonging to the genus '' Hippocrepis'' in the family Fabaceae. Description The overall appearance depends on its habitat: sometimes it forms upright clump ...
,
Lotus corniculatus ''Lotus corniculatus'' is a flowering plant in the pea family Fabaceae. Common names include common bird's-foot trefoil, eggs and bacon, birdsfoot deervetch, and just bird's-foot trefoil (a name also often applied to other ''Lotus'' spp.). It ha ...
,
Securigera varia ''Securigera varia'' (Synonym (taxonomy), synonym ''Coronilla varia''), commonly known as crownvetch or purple crown vetch, is a low-growing legume vine. It is native to Africa, Asia and Europe and is commonly used throughout the United States an ...
, Tetragonalobus maritimus''. Almost all reports relate to observations of oviposition or egg discoveries, only a single detection of an adult caterpillar was found in ''Lotus corniculatus''. In Great Britain eggs were found on the tender young leaves of
bird's-foot trefoil ''Lotus corniculatus'' is a flowering plant in the pea family Fabaceae. Common names include common bird's-foot trefoil, eggs and bacon, birdsfoot deervetch, and just bird's-foot trefoil (a name also often applied to other ''Lotus'' spp.). It ha ...
(''Lotus corniculatus''), the favoured food plant here (although
horseshoe vetch ''Hippocrepis comosa'', the horseshoe vetch, is a species of perennial flowering plant belonging to the genus '' Hippocrepis'' in the family Fabaceae. Description The overall appearance depends on its habitat: sometimes it forms upright clump ...
(''Hippocrepis comosa'') and greater bird's-foot trefoil (''Lotus pendunculatus'') are sometimes used). Other larval host plants in Europe are ''
Eryngium ''Eryngium'' is a genus of flowering plants in the family Apiaceae. There are about 250 species. The genus has a cosmopolitan distribution, with centres of diversity in the western Mediterranean, South America and Mexico.Calviño, C.I., Martíne ...
'', ''
Coronilla The genus ''Coronilla'' contains 8 species of flowering plants native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specif ...
'', ''
Medicago ''Medicago'' is a genus of flowering plants, commonly known as medick or burclover, in the legume family (Fabaceae). It contains at least 87 species and is distributed mainly around the Mediterranean Basin, and extending across temperate Eurasia ...
'', etc. The hemispherical egg with clear longitudinal ribs is initially light yellow and later orange-red in color and is easily recognizable on the green upper side of the leaf with a targeted search. This combination of features results in a further, reliable detection method for the species. During the day, the caterpillar hides in a hiding place made of spun leaves and mainly eats at night. The
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
creates a shelter by spinning leaves together and feeds until fully grown in August. It then creates a larger tent to form a hibernaculum where it
hibernate Hibernation is a state of minimal activity and metabolic reduction entered by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It is most ...
s.
Pupa A pupa (; : pupae) is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages th ...
tion occurs the following spring without further feeding. The winter web is dense like parchment, so that no water can penetrate. Spun threads in summer, the pupae of which result in second generation butterflies, are of a loose texture.


Conservation

In accordance with its preference for low and gappy vegetation, grazing is particularly suitable for maintaining the habitats, which, in addition to the continuous shortening of the vegetation, also results in the formation of the open ground areas that are essential for the species due to the footfall. Extensive grazing with sheep or cattle must therefore be continued on lime lawns, the most important type of habitat in central Europe. If grazing is not possible on poor grass, mowing can also be considered. There are no restrictions with regard to the time of mowing, as the pre-imaginal stages almost always live close to the ground. For the occurrence in poor meadows, the extensive renouncement of fertilization is necessary for survival. Another contribution to the protection of the species is the preservation of small structures with patchy vegetation on fields, embankments and roadsides. It is essential to avoid the use of suction mowers or mulchers when caring for such structures. The creation of new habitats is also possible in sub-areas of the occurrence, for example on limestone gravel in the valleys of southern Bavaria, since the imagos often colonize anthropogenic secondary sites. Suitable areas are e.g. flood dams, embankments or disused mining sites, which are maintained in a low-growing and gappy state by largely dispensing with planting and corresponding subsequent care.


Etymology

"A grandson of Jupiter"


See also

*
List of butterflies of Great Britain This is a list of butterfly, butterflies of Great Britain, including extinct, naturalised species and those of dubious origin. The list comprises butterfly species listed in ''The Moths and Butterflies of Great Britain and Ireland'' by Emmet ''et a ...


References


External links


Butterfly Conservation



UK Butterflies

Butterfly Conservation Armenia
{{Taxonbar, from=Q867291 Erynnis Butterflies of Asia Butterflies of Europe Butterflies described in 1758 Taxa named by Carl Linnaeus