Ergastic substances are non-
protoplasmic materials found in
cells
Cell most often refers to:
* Cell (biology), the functional basic unit of life
Cell may also refer to:
Locations
* Monastic cell, a small room, hut, or cave in which a religious recluse lives, alternatively the small precursor of a monastery w ...
.
The living protoplasm of a cell is sometimes called the bioplasm and distinct from the ergastic substances of the cell. The latter are usually organic or inorganic substances that are products of metabolism, and include crystals, oil drops, gums,
tannin
Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids.
The term ''tannin'' (from Anglo-Norman ''tanner ...
s, resins and other compounds that can aid the organism in defense, maintenance of cellular structure, or just substance storage. Ergastic substances may appear in the
protoplasm
Protoplasm (; ) is the living part of a cell that is surrounded by a plasma membrane. It is a mixture of small molecules such as ions, monosaccharides, amino acid, and macromolecules such as proteins, polysaccharides, lipids, etc.
In some defin ...
, in
vacuole
A vacuole () is a membrane-bound organelle which is present in plant and fungal cells and some protist, animal, and bacterial cells. Vacuoles are essentially enclosed compartments which are filled with water containing inorganic and organic ...
s, or in the
cell wall
A cell wall is a structural layer surrounding some types of cells, just outside the cell membrane. It can be tough, flexible, and sometimes rigid. It provides the cell with both structural support and protection, and also acts as a filtering mec ...
.
Carbohydrates
Reserve carbohydrate of plants are the derivatives of the end products of photosynthesis.
Cellulose
Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wall ...
and
starch
Starch or amylum is a polymeric carbohydrate consisting of numerous glucose units joined by glycosidic bonds. This polysaccharide is produced by most green plants for energy storage. Worldwide, it is the most common carbohydrate in human diets ...
are the main ergastic substances of plant cells. Cellulose is the chief component of the cell wall, and starch occurs as a reserve material in the protoplasm.
Starch, as starch grains, arise almost exclusively in
plastid
The plastid (Greek: πλαστός; plastós: formed, molded – plural plastids) is a membrane-bound organelle found in the cells of plants, algae, and some other eukaryotic organisms. They are considered to be intracellular endosymbiotic cyanobac ...
s, especially
leucoplast
Leucoplasts (λευκός leukós "white", πλαστός plastós "formed, molded") are a category of plastid and as such are organelles found in plant cells. They are non-pigmented, in contrast to other plastids such as the chloroplast.
Lacki ...
s and
amyloplast
Amyloplasts are a type of plastid, double-enveloped organelles in plant cells that are involved in various biological pathways. Amyloplasts are specifically a type of leucoplast, a subcategory for colorless, non-pigment-containing plastids. Amylop ...
s.
Proteins
Although
protein
Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, respon ...
s are the main component of living protoplasm, proteins can occur as inactive, ergastic bodies—in an amorphous or crystalline (or crystalloid) form. A well-known amorphous ergastic protein is
gluten
Gluten is a structural protein naturally found in certain cereal grains. Although "gluten" often only refers to wheat proteins, in medical literature it refers to the combination of prolamin and glutelin proteins naturally occurring in all gra ...
.
Fats and oils
Fat
In nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such compounds, most commonly those that occur in living beings or in food.
The term often refers specifically to triglycerides (triple es ...
s (
lipid
Lipids are a broad group of naturally-occurring molecules which includes fats, waxes, sterols, fat-soluble vitamins (such as vitamins A, D, E and K), monoglycerides, diglycerides, phospholipids, and others. The functions of lipids incl ...
s) and oils are widely distributed in plant tissues. Substances related to fats—
waxes,
suberin
Suberin, cutin and lignins are complex, higher plant epidermis and periderm cell-wall macromolecules, forming a protective barrier. Suberin, a complex polyester biopolymer, is lipophilic, and composed of long chain fatty acids called suberin aci ...
, and
cutin
Cutin is one of two waxy polymers that are the main components of the plant cuticle, which covers all aerial surfaces of plants. It is an insoluble substance with waterproof quality. Cutin also harbors cuticular waxes, which assist in cuticle st ...
—occur as protective layers in or on the cell wall.
Crystals
Animals eliminate excess inorganic materials; plants mostly deposit such material in their tissues. Such
mineral
In geology and mineralogy, a mineral or mineral species is, broadly speaking, a solid chemical compound with a fairly well-defined chemical composition and a specific crystal structure that occurs naturally in pure form.John P. Rafferty, ed. ...
matter is mostly salts of
calcium
Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar t ...
and anhydrides of
silica
Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is o ...
.
*
Raphides
Raphides (pronounced /ˈræfɪˌdiz/, singular raphide /ˈreɪfʌɪd/ or raphis) are needle-shaped crystals of calcium oxalate monohydrate (prismatic monoclinic crystals) or calcium carbonate as aragonite ( dipyramidal orthorhombic crystals), ...
are a type of elongated crystalline form of calcium oxalate aggregated in bundles within a plant cell. Because of the needle-like form, large numbers in the tissue of, say, a leaf can render the leaf unpalatable to herbivores (see
Dieffenbachia
''Dieffenbachia'' , commonly known as dumb cane or leopard lily, is a genus of tropical flowering plants in the family Araceae. It is native to the New World Tropics from Mexico and the West Indies south to Argentina. Some species are widely cul ...
and
taro
Taro () (''Colocasia esculenta)'' is a root vegetable. It is the most widely cultivated species of several plants in the family Araceae that are used as vegetables for their corms, leaves, and petioles. Taro corms are a food staple in Afric ...
).
*
Druse
*
Cystolith
Cystolith (Gr. "cavity" and "stone") is a botanical term for outgrowths of the epidermal cell wall, usually of calcium carbonate, formed in a cellulose matrix in special cells called lithocysts, generally in the leaf of plants.
Cystoliths are ...
References
{{reflist
Cell anatomy