An erection (clinically: penile erection or penile tumescence) is a
physiological
Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemica ...
phenomenon in which the
penis
A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males d ...
becomes firm, engorged, and enlarged. Penile erection is the result of a complex interaction of psychological, neural, vascular, and endocrine factors, and is often associated with
sexual arousal or
sexual attraction
Sexual attraction is attraction on the basis of sexual desire or the quality of arousing such interest. Sexual attractiveness or sex appeal is an individual's ability to attract other people sexually, and is a factor in sexual selection or ma ...
, although erections can also be spontaneous. The shape, angle, and direction of an erection varies considerably between humans.
Physiologically, an erection is required for a male to effect vaginal penetration or
sexual intercourse and is triggered by the
parasympathetic
The parasympathetic nervous system (PSNS) is one of the three divisions of the autonomic nervous system, the others being the sympathetic nervous system and the enteric nervous system. The enteric nervous system is sometimes considered part o ...
division of the
autonomic nervous system, causing the levels of
nitric oxide
Nitric oxide (nitrogen oxide or nitrogen monoxide) is a colorless gas with the formula . It is one of the principal oxides of nitrogen. Nitric oxide is a free radical: it has an unpaired electron, which is sometimes denoted by a dot in its ...
(a
vasodilator
Vasodilation is the widening of blood vessels. It results from relaxation of smooth muscle cells within the vessel walls, in particular in the large veins, large arteries, and smaller arterioles. The process is the opposite of vasoconstriction ...
) to rise in the
trabecular arteries and
smooth muscle
Smooth muscle is an involuntary non- striated muscle, so-called because it has no sarcomeres and therefore no striations (''bands'' or ''stripes''). It is divided into two subgroups, single-unit and multiunit smooth muscle. Within single-unit ...
of the penis. The arteries
dilate causing the
corpora cavernosa of the penis (and to a lesser extent the
corpus spongiosum) to fill with
blood
Blood is a body fluid in the circulatory system of humans and other vertebrates that delivers necessary substances such as nutrients and oxygen to the cells, and transports metabolic waste products away from those same cells. Blood in th ...
; simultaneously the
ischiocavernosus and
bulbospongiosus muscles compress the
vein
Veins are blood vessels in humans and most other animals that carry blood towards the heart. Most veins carry deoxygenated blood from the tissues back to the heart; exceptions are the pulmonary and umbilical veins, both of which carry oxygenate ...
s of the corpora cavernosa restricting the egress and circulation of this blood. Erection subsides when parasympathetic activity reduces to baseline.
As an autonomic nervous system response, an erection may result from a variety of stimuli, including
sexual stimulation
Sexual stimulation is any stimulus (including bodily contact) that leads to, enhances and maintains sexual arousal, and may lead to orgasm. Although sexual arousal may arise without physical stimulation, achieving orgasm usually requires physic ...
and
sexual arousal, and is therefore not entirely under conscious control. Erections during sleep or upon waking up are known as
nocturnal penile tumescence (NPT), also known as "morning wood". Absence of nocturnal erection is commonly used to distinguish between physical and psychological causes of
erectile dysfunction
Erectile dysfunction (ED), also called impotence, is the type of sexual dysfunction in which the penis fails to become or stay erect during sexual activity. It is the most common sexual problem in men.Cunningham GR, Rosen RC. Overview of ma ...
and
impotence.
The state of a penis which is partly, but not fully, erect is sometimes known as semi-erection (clinically: partial tumescence); a penis which is not erect is typically referred to as being flaccid, or soft.
Physiology
An erection is necessary for
natural insemination as well as for the harvesting of sperm for
artificial insemination
Artificial insemination is the deliberate introduction of sperm into a female's cervix or uterine cavity for the purpose of achieving a pregnancy through in vivo fertilization by means other than sexual intercourse. It is a fertility treatment ...
, are common for children and infants, and even occur before birth. After reaching
puberty
Puberty is the process of physical changes through which a child's body matures into an adult body capable of sexual reproduction. It is initiated by hormonal signals from the brain to the gonads: the ovaries in a girl, the testes in a b ...
, erections occur much more frequently.
it occurs when two tubular structures, called the
corpora cavernosa, that run the length of the penis, become engorged with
venous blood. This may result from any of various
physiological
Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemica ...
stimuli, also known as
sexual stimulation
Sexual stimulation is any stimulus (including bodily contact) that leads to, enhances and maintains sexual arousal, and may lead to orgasm. Although sexual arousal may arise without physical stimulation, achieving orgasm usually requires physic ...
and
sexual arousal. The
corpus spongiosum is a single tubular structure located just below the corpora cavernosa, which contains the
urethra
The urethra (from Greek οὐρήθρα – ''ourḗthrā'') is a tube that connects the urinary bladder to the urinary meatus for the removal of urine from the body of both females and males. In human females and other primates, the urethra c ...
, through which
urine
Urine is a liquid by-product of metabolism in humans and in many other animals. Urine flows from the kidneys through the ureters to the urinary bladder. Urination results in urine being excreted from the body through the urethra.
Cellular ...
and
semen
Semen, also known as seminal fluid, is an organic bodily fluid created to contain spermatozoa. It is secreted by the gonads (sexual glands) and other sexual organs of male or hermaphroditic animals and can fertilize the female ovum. Semen ...
pass during
urination
Urination, also known as micturition, is the release of urine from the urinary bladder through the urethra to the outside of the body. It is the urinary system's form of excretion. It is also known medically as micturition, voiding, ure ...
and
ejaculation
Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the male reproductory tract as a result of an orgasm. It is the final stage and natural objective of male sexual stimulation, and an essential compone ...
respectively. This may also become slightly engorged with blood, but less so than the corpora cavernosa.
The
scrotum
The scrotum or scrotal sac is an anatomical male reproductive structure located at the base of the penis that consists of a suspended dual-chambered sac of skin and smooth muscle. It is present in most terrestrial male mammals. The scrotum co ...
may, but does not always, become tightened during erection. Generally, in uncircumcised males, the
foreskin
In male human anatomy, the foreskin, also known as the prepuce, is the double-layered fold of skin, mucosal and muscular tissue at the distal end of the human penis that covers the glans and the urinary meatus. The foreskin is attached to ...
automatically and gradually retracts, exposing the
glans, though some people may have to manually retract their foreskin.
Autonomic control
In the presence of mechanical stimulation, erection is initiated by the
parasympathetic
The parasympathetic nervous system (PSNS) is one of the three divisions of the autonomic nervous system, the others being the sympathetic nervous system and the enteric nervous system. The enteric nervous system is sometimes considered part o ...
division of the
autonomic nervous system with minimal input from the
central nervous system
The central nervous system (CNS) is the part of the nervous system consisting primarily of the brain and spinal cord. The CNS is so named because the brain integrates the received information and coordinates and influences the activity of all p ...
. Parasympathetic branches extend from the
sacral plexus
In human anatomy, the sacral plexus is a nerve plexus which provides motor and sensory nerves for the posterior thigh, most of the lower leg and foot, and part of the pelvis. It is part of the lumbosacral plexus and emerges from the lumbar ver ...
into the
arteries supplying the erectile tissue; upon stimulation, these nerve branches release
acetylcholine
Acetylcholine (ACh) is an organic chemical that functions in the brain and body of many types of animals (including humans) as a neurotransmitter. Its name is derived from its chemical structure: it is an ester of acetic acid and choline. Par ...
, which in turn causes release of
nitric oxide
Nitric oxide (nitrogen oxide or nitrogen monoxide) is a colorless gas with the formula . It is one of the principal oxides of nitrogen. Nitric oxide is a free radical: it has an unpaired electron, which is sometimes denoted by a dot in its ...
from
endothelial cell
The endothelium is a single layer of squamous endothelial cells that line the interior surface of blood vessels and lymphatic vessels. The endothelium forms an interface between circulating blood or lymph in the lumen and the rest of the ves ...
s in the
trabecular arteries. Nitric oxide diffuses to the
smooth muscle
Smooth muscle is an involuntary non- striated muscle, so-called because it has no sarcomeres and therefore no striations (''bands'' or ''stripes''). It is divided into two subgroups, single-unit and multiunit smooth muscle. Within single-unit ...
of the arteries (called ''trabecular smooth muscle''), acting as a
vasodilating agent.
The arteries dilate, filling the ''corpus spongiosum'' and ''corpora cavernosa'' with blood. The
ischiocavernosus and
bulbospongiosus muscles also compress the veins of the corpora cavernosa, limiting the venous drainage of blood. Erection subsides when parasympathetic stimulation is discontinued; baseline stimulation from the
sympathetic division of the autonomic nervous system causes constriction of the penile arteries and cavernosal sinosoids, forcing blood out of the erectile tissue via erection-related veins which include one deep dorsal vein, a pair of cavernosal veins, and two pairs of para-arterial veins between Buck's fascia and the tunica albuginea.
Erection rigidity is mechanically controlled by reduction blood flow via theses veins, and thereby building up the pressure of the corpus cavernosum and corpus spongiosum, an integral instructure, the distal ligament, buttresses the glans penis.
After
ejaculation
Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the male reproductory tract as a result of an orgasm. It is the final stage and natural objective of male sexual stimulation, and an essential compone ...
or cessation of stimulation, erection usually subsides, but the time taken may vary depending on the length and thickness of the penis.
Voluntary and involuntary control
The
cerebral cortex
The cerebral cortex, also known as the cerebral mantle, is the outer layer of neural tissue of the cerebrum of the brain in humans and other mammals. The cerebral cortex mostly consists of the six-layered neocortex, with just 10% consisting o ...
can initiate erection in the absence of direct mechanical stimulation (in response to visual, auditory, olfactory, imagined, or tactile stimuli) acting through erectile centers in the lumbar and sacral regions of the
spinal cord
The spinal cord is a long, thin, tubular structure made up of nervous tissue, which extends from the medulla oblongata in the brainstem to the lumbar region of the vertebral column (backbone). The backbone encloses the central canal of the spin ...
. The cortex may suppress erection, even in the presence of mechanical stimulation, as may other psychological, emotional, and environmental factors.
Nocturnal erection
The penis may become erect during sleep or be erect on waking up. Such an erection is medically known as
nocturnal penile tumescence (informally: ''morning wood'' or ''morning glory'').
Urination
Urinating with an erection can be more difficult than without. David Samadi, director of men's health at
St. Francis Hospital in
Long Island, says "To prevent
semen
Semen, also known as seminal fluid, is an organic bodily fluid created to contain spermatozoa. It is secreted by the gonads (sexual glands) and other sexual organs of male or hermaphroditic animals and can fertilize the female ovum. Semen ...
from entering into the
bladder, the
internal urethral sphincter
The internal urethral sphincter is a urethral sphincter muscle which constricts the internal urethral orifice. It is located at the junction of the urethra with the urinary bladder and is continuous with the detrusor muscle, but anatomically and ...
contracts."
This also works in opposite way, making it less likely that urine will pass through the urethra with an erect penis. Combined with
morning wood, and a full bladder from overnight, many people may have experience with the extra challenge that comes with the combination.
Medical recommendations include:
# Waiting for the erection to go away, including techniques for relaxing
# Peeing while bending over or sitting down
# Massaging, or putting light pressure or a hot compress on the bladder (from the outside, between the navel and the pubic bone)
[
]
Socio-sexual aspects
Social
Though an erection can have many causes, the more common indicator of sexual arousal has a prevalent view over society and its portrayal in public is considered taboo in many societies, probably lesser than public sex
__NOTOC__
Public sex is sexual activity that takes place in a public context. It refers to one or more persons performing a sex act in a public place, or in a private place that can be viewed from a public place.
Such a private place may be ...
but higher than nudity
Nudity is the state of being in which a human is without clothing.
The loss of body hair was one of the physical characteristics that marked the biological evolution of modern humans from their hominin ancestors. Adaptations related to h ...
. Erectile dysfunction is considered a flaw that associates shame at the individual and his partner.
The penile plethysmograph, which measures erections, has been used by some governments and courts of law to measure sexual orientation. An unusual aversion to the erect penis is sometimes referred to as phallophobia.
Spontaneous or random erections
Spontaneous erections, also known as involuntary, random or unwanted erections, are commonplace and a normal part of male physiology. Socially, such erections can be embarrassing
Embarrassment or awkwardness is an emotional state that is associated with mild to severe levels of discomfort, and which is usually experienced when someone commits (or thinks of) a socially unacceptable or frowned-upon act that is witnessed ...
if they happen in public or when undesired. Such erections can occur at any time of day, and if clothed may cause a bulge which (if required) can be disguised or hidden by wearing close-fitting underwear, a long shirt, or baggier clothes. Few slang terms associated with erect penis outline becoming visible from underneath clothing are ''mooseknuckle'' and ''manbulge''.
Size
The length of the flaccid penis is not necessarily indicative of the length of the penis when it becomes erect, with some smaller flaccid penises growing much longer, and some larger flaccid penises growing comparatively less.
Generally, the size of an erect penis is fixed throughout post- pubescent life. Its size may be increased by surgery, although penile enlargement is controversial, and a majority of men were "not satisfied" with the results, according to one study.
Though the size of a penis varies considerably between males, the average length of an erect human penis
The human penis is an external male intromittent organ that additionally serves as the urinary duct. The main parts are the root (radix); the body (corpus); and the epithelium of the penis including the shaft skin and the foreskin (prepuce) ...
is 13.12 cm (5.17 inches), while the average circumference of an erect human penis is 11.66 cm (4.59 inches).
Direction
Although many erect penises point upwards, it is common and normal for the erect penis to point nearly vertically upwards or horizontally straight forward or even nearly vertically downwards, all depending on the tension of the suspensory ligament that holds it in position. An erect penis can also take on a number of different shapes, ranging from a straight tube to a tube with a curvature up or down or to the left or right. An increase in penile curvature can be caused by Peyronie's disease. This may cause physical and psychological effects for the affected individual, which could include erectile dysfunction
Erectile dysfunction (ED), also called impotence, is the type of sexual dysfunction in which the penis fails to become or stay erect during sexual activity. It is the most common sexual problem in men.Cunningham GR, Rosen RC. Overview of ma ...
or pain during an erection. Treatments include oral medication (such as colchicine
Colchicine is a medication used to treat gout and Behçet's disease. In gout, it is less preferred to NSAIDs or steroids. Other uses for colchicine include the management of pericarditis and familial Mediterranean fever. Colchicine is tak ...
) or surgery, which is most often performed only as a last resort.
The following table shows how common various erection angles are for a standing male. In the table, zero degrees (0°) is pointing straight up against the abdomen, 90° is horizontal and pointing straight forward, and 180° is pointing straight down to the feet. An upward pointing angle is most common and the average erection angle is 74.3 degrees. The penile curvature was measured same time. 63% men have straight penis. 22.2% men have upwards curvature and 14.8% men have downwards curvature.
Medical conditions
Erectile dysfunction
Erectile dysfunction (also known as ED or "(male) impotence") is a sexual dysfunction
Sexual dysfunction is difficulty experienced by an individual or partners during any stage of normal sexual activity, including physical pleasure, desire, preference, arousal, or orgasm. The World Health Organization defines sexual dysfunction ...
characterized by the inability to develop and/or maintain an erection. The study of erectile dysfunction within medicine is known as andrology
Andrology (from grc, ἀνήρ, ''anēr'', genitive , ''andros'', "man"; and , '' -logia'') is a name for the medical specialty that deals with male health, particularly relating to the problems of the male reproductive system and urological p ...
, a sub-field within urology
Urology (from Greek οὖρον ''ouron'' "urine" and '' -logia'' "study of"), also known as genitourinary surgery, is the branch of medicine that focuses on surgical and medical diseases of the urinary-tract system and the reproductive org ...
.
Erectile dysfunction may occur due to physiological
Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemica ...
or psychological
Psychology is the scientific study of mind and behavior. Psychology includes the study of conscious and unconscious phenomena, including feelings and thoughts. It is an academic discipline of immense scope, crossing the boundaries betw ...
reasons, most of which are amenable to treatment. Common physiological reasons include diabetes, kidney disease, chronic alcoholism, multiple sclerosis, atherosclerosis, vascular disease, including arterial insufficiency and venogenic erectile dysfunction, and neurologic disease which collectively account for about 70% of ED cases.[ Some drugs used to treat other conditions, such as ]lithium
Lithium (from el, λίθος, lithos, lit=stone) is a chemical element with the symbol Li and atomic number 3. It is a soft, silvery-white alkali metal. Under standard conditions, it is the least dense metal and the least dense solid ...
and paroxetine, may cause erectile dysfunction.
Erectile dysfunction, tied closely as it is to cultural notions of potency, success
Success is the state or condition of meeting a defined range of expectations. It may be viewed as the opposite of failure. The criteria for success depend on context, and may be relative to a particular observer or belief system. One person mig ...
and masculinity
Masculinity (also called manhood or manliness) is a set of attributes, behaviors, and roles associated with men and boys. Masculinity can be theoretically understood as socially constructed, and there is also evidence that some behaviors ...
, can have devastating psychological consequences including feelings of shame
Shame is an unpleasant self-conscious emotion often associated with negative self-evaluation; motivation to quit; and feelings of pain, exposure, distrust, powerlessness, and worthlessness.
Definition
Shame is a discrete, basic emotion, d ...
, loss or inadequacy. There is a strong culture of silence and inability to discuss the matter. Around one in ten men experience recurring impotence problems at some point in their lives.
Priapism
Priapism is a painful condition in which the penis does not return to its flaccid state, despite the absence of both physical and psychological stimulation. Priapism lasting over four hours is a medical emergency.
In non-human animals
At the time of penetration, the canine penis is not erect, and only able to penetrate the female because it includes a narrow bone called the baculum, a feature of most placental mammals. After the male achieves penetration, he will often hold the female tighter and thrust faster, and it is during this time that the male's penis expands. Unlike human sexual intercourse, where the male penis commonly becomes erect before entering the female, canine copulation involves the male first penetrating the female, after which swelling of the penis to erection occurs.
An elephant's penis is S-shaped when fully erect and has a Y-shaped orifice.
Given the small amount of erectile tissue in a bull's penis, there is little enlargement after erection. The penis is quite rigid when non-erect, and becomes even more rigid during erection. Protrusion is not affected much by erection, but more by relaxation of the retractor penis muscle and straightening of the sigmoid flexure.
A male fossa's penis
A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males d ...
reaches to between his forelegs when erect.
When not erect, a horse's penis is housed within the prepuce, long and in diameter with the distal end . The retractor muscle
Retractor may refer to:
* Retractor (medical), a medical instrument
* Retractor (memory), a person
* Retractor (chess), type of a chess problem
* Retractor muscle
Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate ...
contracts to retract the penis into the sheath and relaxes to allow the penis to extend from the sheath. When erect, the penis doubles in length and thickness and the glans increases by 3 to . Erection and protrusion take place gradually, by the increasing tumescence of the erectile vascular tissue in the '' corpus cavernosum penis''. Most stallions achieve erection within 2 minutes of contact with an estrus mare, and mount the estrus mare 5–10 seconds afterward.
A bird penis is different in structure from mammal penises, being an erectile expansion of the cloacal wall and being erected by lymph
Lymph (from Latin, , meaning "water") is the fluid that flows through the lymphatic system, a system composed of lymph vessels (channels) and intervening lymph nodes whose function, like the venous system, is to return fluid from the tissues ...
, not blood. The penis of the lake duck
The lake duck (''Oxyura vittata'') is a small, South American stiff-tailed duck. It is also called the Argentine blue-bill, Argentine blue-billed duck, Argentine lake duck, or Argentine ruddy duck.
Description
The lake duck grows to . Females ...
can reach about the same length as the animal himself when fully erect, but more commonly is about half the bird's length.
Terminology
Clinically, erection is often known as "penile erection", and the state of being erect, and process of erection, are described as "tumescence" or "penile tumescence". The term for the subsiding or cessation of an erection is " detumescence".
Colloquially and in slang
Slang is vocabulary (words, phrases, and usage (language), linguistic usages) of an informal register, common in spoken conversation but avoided in formal writing. It also sometimes refers to the language generally exclusive to the members of p ...
, erection is known by many informal terms. Commonly encountered English terms include 'stiffy', 'hard-on', 'boner' and 'woody'. There are several slang words, euphemisms and synonyms for an erection in English and in other languages. (See also The WikiSaurus entry.)
See also
* Clitoral erection
* Cock ring
* Death erection
*Human penis
The human penis is an external male intromittent organ that additionally serves as the urinary duct. The main parts are the root (radix); the body (corpus); and the epithelium of the penis including the shaft skin and the foreskin (prepuce) ...
* Issues in social nudity
* Nipple stimulation
* Nocturnal penile tumescence
* Priapism
*Sexual function Sexual function is how the body reacts in different stages of the sexual response cycle, or as a result of sexual dysfunction. Relevant aspects of sexual function are defined on the basis of a modified version of Masters and Johnson's work. The asp ...
References
{{Authority control
Sexual arousal
Andrology
Men's health