Equus Altidens
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''Equus altidens'' is an extinct species of equine native to western Eurasia including Europe during the
Early Pleistocene The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...
to early
Middle Pleistocene The Chibanian, more widely known as the Middle Pleistocene (its previous informal name), is an Age (geology), age in the international geologic timescale or a Stage (stratigraphy), stage in chronostratigraphy, being a division of the Pleistocen ...
.


Taxonomy

''Equus altidens'' was first described in 1915 from remains at the early
Middle Pleistocene The Chibanian, more widely known as the Middle Pleistocene (its previous informal name), is an Age (geology), age in the international geologic timescale or a Stage (stratigraphy), stage in chronostratigraphy, being a division of the Pleistocen ...
Sussenborn locality in Germany. Most recent authors consider the species ''Equus marxi'' also known from Sussenborn to be a
junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...
of ''E. altidens''. The majority of authors have suggested the species is related to other "stenonine" equines known from Early Pleistocene of Europe such as ''
Equus stenonis ''Equus stenonis'' is an extinct species of equine that lived in Western Eurasia including Europe during the Early Pleistocene. Taxonomy and evolution The species was first named in 1867, with the type specimen being IGF 560, a skull with a n ...
,'' which are thought to be more closely related to living
zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: Grévy's zebra (''Equus grevyi''), the plains zebra (''E. quagga''), and the mountain zebra (''E. ...
s and
asses Ass most commonly refers to: * Buttocks (in informal American English) * Donkey or ass, ''Equus africanus asinus'' **any other member of the subgenus ''Asinus'' Ass or ASS may also refer to: Art and entertainment * ''Ass'' (album), 1973 alb ...
than to true horses. Some authors have suggested a closer relationship to asses than to zebras for ''E. altidens''. Some authors have placed the species in the subgenus ''Sussemionus,'' or place it with other "stenonines" in the separate genus ''Allohippus.'' " Orce Man", a supposed archaic human fossil from the Early Pleistocene of Spain, was later determined to actually be the skull remains of a juvenile of this species. However, a later discovery of a tooth confirmed the presence of archaic humans at the same site.


Description

''Equus altidens'' is a medium-sized equine species with an estimated body mass of around .ST2 The species had slender limbs, the most slender among the "stenonines", including the
metapodial Metapodials are long bone The long bones are those that are longer than they are wide. They are one of five types of bones: long, short, flat, irregular and sesamoid. Long bones, especially the femur and tibia, are subjected to most of the l ...
bones, and is distinguished from other equines by a number of dental morphological features, including narrow and small crowns, and deep molar ectoflexid, though the dental morphology exhibits considerable variability in a number of characters.


Distribution and chronology

''Equus altidens is'' known from remains found across Europe, spanning from the Iberian Peninsula to Italy and Greece, northwards to Germany and Britain, and as far eastwards as Georgia in the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
. Other possible records are known from Romania, Turkey, Tajikistan, Hungary, and Russia. Authors have differed about the timing of the earliest appearance of the species, ranging from as late as 1.2 million years ago to as early as 1.8 million years ago, with recent research supporting the earlier end of the range, with the earliest records coming from the Dmanisi site in Georgia. The youngest remains of the species date to around 600,000 years ago, when the species like other European "stenonine" equines was replaced by early caballine true horses belonging to the species '' Equus mosbachensis''.


Ecology

Dental wear analysis that ''Equus altidens'' had an abrasive, largely grazing based diet, though it may have seasonally engaged in mixed feeding. It often co-occurred alongside another larger "stenonine" equine, ''
Equus suessenbornensis ''Equus suessenbornensis'' is an extinct species of large equine native to Western Eurasia, including Europe, during the Early Pleistocene to early Middle Pleistocene. Taxonomy The species was first described in 1900 based on remains found at the ...
.'' Evidence suggests that this species had a diet that consisted of considerably more browse than ''E. altidens'', suggesting dietary
niche partitioning In ecology, a niche is the match of a species to a specific environmental condition. Three variants of ecological niche are described by It describes how an organism or population responds to the distribution of resources and competitors (for e ...
between the two species. ''Equus altidens'' is thought to have primarily inhabited arid, open habitats. Isotopic analysis from the Venta Micena locality in southeast Spain dating to the Early Pleistocene, around 1.6 million years ago, suggests that at this locality ''Equus altidens'' was regularly preyed upon by sabertooth cats, including the lion sized ''
Homotherium ''Homotherium'' is an extinct genus of Homotherini, scimitar-toothed cat belonging to the extinct subfamily Machairodontinae that inhabited North America, Eurasia, and Africa, as well as possibly South America during the Pliocene and Pleistocene ...
'', and the smaller, jaguar sized ''
Megantereon ''Megantereon'' is an extinct genus of prehistoric machairodontine saber-toothed cat that lived in Eurasia, Africa and possibly North America from the late Pliocene to the Middle Pleistocene. It is a member of the tribe Smilodontini, and close ...
''.


Relationship with humans

At the Fuente Nueva-3 site in Spain, during the late Early Pleistocene around 1.2 million years ago, cut marks found on bones indicate butchery of ''Equus altidens'' by archaic humans, likely ''
Homo antecessor ''Homo antecessor'' (Latin "pioneer man") is an extinct species of archaic human recorded in the Spanish Archaeological Site of Atapuerca, Sierra de Atapuerca, a productive archaeological site, from 1.2 to 0.8 million years ago during the Early ...
''.


References

{{Taxonbar, from1=Q109313408 altidens Prehistoric Perissodactyla