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''Equisetum telmateia'', the great horsetail, is a species of ''
Equisetum ''Equisetum'' (; horsetail) is the only living genus in Equisetaceae, a family of vascular plants that reproduce by spores rather than seeds. ''Equisetum'' is a "living fossil", the only living genus of the entire subclass Equisetidae, which ...
'' (horsetail)
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
to
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
, western
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
and northwest
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
.Hyde, H. A., Wade, A. E., & Harrison, S. G. (1978). ''Welsh Ferns''. National Museum of Wales . It was formerly widely treated in a broader sense including a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
(subsp. ''braunii'') in western North America, but this is now treated as a separate species, '' Equisetum braunii''.


Description

It is a
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of ...
perennial plant In horticulture, the term perennial (''wikt:per-#Prefix, per-'' + ''wikt:-ennial#Suffix, -ennial'', "through the year") is used to differentiate a plant from shorter-lived annual plant, annuals and biennial plant, biennials. It has thus been d ...
, with separate green
photosynthetic Photosynthesis ( ) is a Biological system, system of biological processes by which Photoautotrophism, photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical ener ...
sterile stems, and pale yellowish non-photosynthetic
spore In biology, a spore is a unit of sexual reproduction, sexual (in fungi) or asexual reproduction that may be adapted for biological dispersal, dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores fo ...
-bearing fertile stems. The sterile stems, produced in late spring and dying down in late autumn, are (rarely to ) tall (the tallest species of horsetail outside of tropical regions) and diameter, heavily branched, with whorls of 14–40 branches, these up to long, diameter and unbranched, emerging from the axils of a ring of
bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...
s; the main stem itself is whitish, without chlorophyll or stomata. The fertile stems are produced in early spring before the sterile shoots, growing to tall with an apical spore-bearing
strobilus A strobilus (: strobili) is a structure present on many land plant species consisting of sporangia-bearing structures densely aggregated along a stem. Strobili are often called cones, but some botanists restrict the use of the term cone to the woo ...
long and broad, and no side branches. The spores disperse in mid spring, with the fertile stems dying immediately after spore release. It also spreads by means of rhizomes that have been observed to penetrate into wet clay soil, spreading laterally in multiple layers. Occasionally plants produce stems that are both fertile and photosynthetic.


Distribution

Great horsetail is widespread across most of Europe north to 56° to 58° N. In
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
and
Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
, it is widespread and often common in suitable habitats, but scarcer in
Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
, particularly north of the
Central Belt The Central Belt of Scotland is the Demographics of Scotland, area of highest population density within Scotland. Depending on the definition used, it has a population of between 2.4 and 4.2 million (the country's total was around 5.4 million in ...
, except for a notable concentration on
Skye The Isle of Skye, or simply Skye, is the largest and northernmost of the major islands in the Inner Hebrides of Scotland. The island's peninsulas radiate from a mountainous hub dominated by the Cuillin, the rocky slopes of which provide some o ...
; the northernmost population of the species anywhere is near
Thurso Thurso (pronounced ; , ) is a town and former burgh on the north coast of the Highland council area of Scotland. Situated in the historical County of Caithness, it is the northernmost town on the island of Great Britain. From a latitudinal s ...
on the north coast of Scotland at about 58°35' N. Further east, it does not reach so far north; in
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
, it is rare and endangered, confined to just three locations north to about 56°08' N on the west coast of the southern province of
Skåne Scania ( ), also known by its native name of Skåne (), is the southernmost of the historical provinces of Sweden, provinces () of Sweden. Located in the south tip of the geographical region of Götaland, the province is roughly conterminous w ...
. Its southern and eastern limits are less well mapped, but it occurs south to
Madeira Madeira ( ; ), officially the Autonomous Region of Madeira (), is an autonomous Regions of Portugal, autonomous region of Portugal. It is an archipelago situated in the North Atlantic Ocean, in the region of Macaronesia, just under north of ...
(32°45' N), the
Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. They separate the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range, which stretches around through M ...
in northwest Africa, and
Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...
, and east to
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
; the easternmost mapped on
iNaturalist iNaturalist is an American 501(c)(3) nonprofit social network of naturalists, citizen scientists, and biologists built on the concept of mapping and sharing observations of biodiversity across the globe. iNaturalist may be accessed via its web ...
('research grade' records only) is at 52°58' E in Iran.


Habitat

It is found in damp shady places, spring
fen A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetland along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires ...
s and
seepage In soil mechanics, seepage is the movement of water through soil. If fluid pressures in a soil deposit are uniformly increasing with depth according to u = \rho_w g z_w, where z_w is the depth below the water table, then hydrostatic conditions wi ...
lines, usually in open woodlands, commonly forming large
clonal colonies A clonal colony or genet is a group of genetically identical individuals, such as plants, fungi, or bacteria, that have grown in a given location, all originating vegetatively, not sexually, from a single ancestor. In plants, an individual in ...
.


References

{{Taxonbar, from=Q936546 telmateia Flora of Europe Flora of Africa Plants described in 1783 Taxa named by Jakob Friedrich Ehrhart Flora of Lebanon