''Epipompilus'' is a genus of spider wasps in the subfamily
Pepsinae, part of the widespread family
Pompilidae. Representatives of ''Epipompilus'' can be found in Australasia and North and South America.
This distribution may indicate that ''Epipompilus'' evolved in
Gondwana
Gondwana ( ; ) was a large landmass, sometimes referred to as a supercontinent. The remnants of Gondwana make up around two-thirds of today's continental area, including South America, Africa, Antarctica, Australia (continent), Australia, Zea ...
and is similar to other Gondwanan
taxa
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
such as the southern beech ''
Nothofagus
''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere, found across southern South America (Chile, Argentina) and east and southeast Australia, New Zealand, New Guin ...
'' and ''
Auracaria''.
''Epipompilus'' is found in North and South America, ranging from
Argentina
Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
to extreme southern
United States
The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
,
with around a dozen known species. One species, ''
E. insularis'' is endemic to
New Zealand
New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
. In Australia, the genus reaches its greatest diversity, with a greater number of species and a more varied spectrum of morphological features than among the American species. The ''Epipompilus'' species in
New Guinea
New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
are notably brilliantly coloured and apparently highly evolved species. The genus is restricted to these areas but several
Tertiary
Tertiary (from Latin, meaning 'third' or 'of the third degree/order..') may refer to:
* Tertiary period, an obsolete geologic period spanning from 66 to 2.6 million years ago
* Tertiary (chemistry), a term describing bonding patterns in organic ch ...
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s from the northern hemisphere should probably be placed in ''Epipompilus''.
Ecology and behaviour
These wasps are scarce in collections, probably due to their small size and the fact that they rarely visit flowers. A single male ''
E. turneri'' which was collected on ''
Leptospermum
''Leptospermum'' is a genus of shrubs and small trees in the myrtle family Myrtaceae commonly known as tea trees, although this name is sometimes also used for some species of ''Melaleuca''. Most species are endemic to Australia, with the greate ...
'' in
New South Wales
New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...
is one of the few flower records. In Australia observation and collection have often been associated with the trunks of living ''
Eucalyptus
''Eucalyptus'' () is a genus of more than 700 species of flowering plants in the family Myrtaceae. Most species of ''Eucalyptus'' are trees, often Mallee (habit), mallees, and a few are shrubs. Along with several other genera in the tribe Eucalyp ...
'' trees. The morphology of many of the species suggests that they are adapted for crawling under bark and for entering crevices to search for spiders. Prey recorded includes spiders from the family
Sparassidae. These wasps probably do not build nests but hunt spiders underneath bark and lay eggs on them as they find them. Taken into consideration with the many primitive structural features of members of this genus, it is suggestive that the hunting technique of ''Epipompilus'' represents an ancestral type of behaviour for spider wasps.
One of three new species identified in 2020,
[ based on a single specimen seen in 2018, '' E. namadji'', is named after the ]Namadgi National Park
Namadgi National Park is a protected area in the southwest of the Australian Capital Territory (ACT), bordering Kosciuszko National Park in New South Wales. It lies approximately southwest of Canberra, and occupies approximately 46 percent of th ...
in the Australian Capital Territory
The Australian Capital Territory (ACT), known as the Federal Capital Territory until 1938, is an internal States and territories of Australia, territory of Australia. Canberra, the capital city of Australia, is situated within the territory, an ...
, in which it was found. Efforts are ongoing by teams from the Australian National Insect Collection at the CSIRO
The Commonwealth Scientific and Industrial Research Organisation (CSIRO) is an Australian Government agency that is responsible for scientific research and its commercial and industrial applications.
CSIRO works with leading organisations arou ...
to find more of the wasps, after nearly 80 per cent of the national park
A national park is a nature park designated for conservation (ethic), conservation purposes because of unparalleled national natural, historic, or cultural significance. It is an area of natural, semi-natural, or developed land that is protecte ...
was lost in the 2019–20 Australian bushfire season
The 201920 Australian bushfire season commenced with serious uncontrolled fires in June 2019.
, fires this season have burned an estimated , destroyed over 5,900 buildings (including 2,779 homes) and killed at least 34 people. An estimated o ...
.
Species
The following species have been assigned to ''Epipompilus'':
*'' Epipompilus albofasciatus'' Evans, 1972
*'' Epipompilus aztecus'' Cresson, 1869
*'' Epipompilus caeruleus'' Evans, 1972
*'' Epipompilus cardaleae'' Evans, 1972
*'' Epipompilus compactus'' Evans, 1972
*'' Epipompilus delicatus'' Turner, 1910
*'' Epipompilus depressus'' Evans, 1962
*'' Epipompilus excelsus'' Bradley, 1944
*'' Epipompilus exleyae'' Evans, 1972
*'' Epipompilus formosus'' Evans, 1972
*'' Epipompilus gilesi'' Turner, 1910
*'' Epipompilus hackeri'' Evans, 1972
*'' Epipompilus inca'' Evans, 1968
*'' Epipompilus incompletus'' Evans, 1972
*'' Epipompilus insularis'' Kohl, 1884
*'' Epipompilus jocosus'' Evans, 1968
*'' Epipompilus matthewsi'' Evans, 1972
*'' Epipompilus mirabundus'' Yuan & Rodriguez, 2020
*'' Epipompilus montivagus'' Evans, 1972
*'' Epipompilus multifasciatus'' Evans, 1972
*'' Epipompilus namadji'' Yuan & Rodriguez, 2020
*'' Epipompilus neboissi'' Evans, 1972
*'' Epipompilus nigribasis'' Banks, 1925
*'' Epipompilus pallidus'' Evans, 1962
*'' Epipompilus papuensis'' Evans, 1972
*'' Epipompilus pauper'' Evans, 1972
*'' Epipompilus pictipennis'' Evans, 1962
*'' Epipompilus pulcherrimus'' Evans, 1955
*'' Epipompilus rieki'' Evans, 1972
*'' Epipompilus submetallicus'' Evans, 1972
*'' Epipompilus taree'' Yuan & Rodriguez, 2020
*'' Epipompilus tasmanicus'' Evans, 1972
*'' Epipompilus tucumanus'' Evans, 1968
*'' Epipompilus turneri'' Evans, 1962
*'' Epipompilus variegatus'' Evans, 1972
References
{{Taxonbar, from=Q5383184
Hymenoptera genera
Pepsinae