''Eomorphippus'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

of notohippid notoungulate that lived from the Late

Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', " ...

to the Early

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...

in what is today

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...

Description

This animal was approximately the size of a

goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of ...

, with a build probably more robust. It had a heavy and sturdy skull, with a relatively short muzzle. The dentition was characterized by high-crowned (hypsodont) incisors, posterior premolars and molars. The teeth didn't have

cementum Cementum is a specialized calcified substance covering the root of a tooth. The cementum is the part of the periodontium that attaches the teeth to the alveolar bone by anchoring the periodontal ligament.Illustrated Dental Embryology, Histology, ...

. The upper incisors were slightly protruding forward. The third upper incisor was wider than the other two, while the fourth premolar was more molariform than the other premolars, but devoid of a distinct hypocone. The upper molars had hypocones, with a variable but generally deep fissure separating them from the protocones. The fissure was blocked by the medial projection of a hooked structure. Apart from the permanent major fossa, the other fossae are erased with wear.

Classification

The genus ''Eomorphippus'' was first described in 1901 by

Florentino Ameghino Florentino Ameghino (born Giovanni Battista Fiorino Giuseppe Ameghino September 19, 1853 – August 6, 1911) was an Argentine naturalist, paleontologist, anthropologist and zoologist, whose fossil discoveries on the Argentine Pampas, especiall ...

, based on fossil remains found in

Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and ...

, in terrains dated either from the Late Eocene or from the Early Oligocene. The type species is ''Eomorphippus obscurus'', whose fossils have also been found in the

Chubut Province Chubut ( es, Provincia del Chubut, ; cy, Talaith Chubut) is a province in southern Argentina, situated between the 42nd parallel south (the border with Río Negro Province), the 46th parallel south (bordering Santa Cruz Province), the Andes ran ...

. Two other species, ''E. bondi'' and ''E. neilopdykei'', were both described in 2018, from remains found in Oligocene terrains from

Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...

. Another species, traditionally attributed to ''Eomorphippus'', ''E. pascuali'', was reassigned in the same 2018 study to the genus '' Rosendo''. ''Eomorphippus'' was a toxodont, a clade of notoungulates, that developed a remarkable diversity of forms during the

Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...

. ''Eomorphippus'' has been historically included within the family

Notohippidae Notohippidae is a paraphyletic extinct family of notoungulate mammals from South America. Notohippids are known from the Eocene and Oligocene epochs.McKenna, Malcolm C., and Bell, Susan K. 1997. ''Classification of Mammals Above the Species Leve ...

, which included several genera whose hypsodont teeth were similar to those of

horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million ...

s. This family, however, is no longer considered

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...

Bibliography

*F. Ameghino. 1901. Notices préliminaires sur des ongulés nouveaux des terrains crétacés de Patagonie reliminary notes on new ungulates from the Cretaceous terrains of Patagonia Boletin de la Academia Nacional de Ciencias de Córdoba 16:349-429 *G. G. Simpson. 1967. The beginning of the age of mammals in South America. Part II. Bulletin of the American Museum of Natural History 137:1-260 {{Taxonbar, from=Q19594762 Toxodonts Eocene mammals of South America Oligocene mammals of South America Paleogene Argentina Fossils of Argentina Paleogene Chile Fossils of Chile Fossil taxa described in 1901 Taxa named by Florentino Ameghino Prehistoric placental genera Golfo San Jorge Basin Sarmiento Formation