Australidelphia is the

superorder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...

that contains roughly three-quarters of all

marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...

s, including all those native to

Australasia Australasia is a region that comprises Australia, New Zealand and some neighbouring islands in the Pacific Ocean. The term is used in a number of different contexts, including geopolitically, physiogeographically, philologically, and ecolo ...

and a single species — the

monito del monte The monito del monte or colocolo opossum, ''Dromiciops gliroides'', also called ''chumaihuén'' in Mapudungun, is a diminutive marsupial native only to southwestern South America (Argentina and Chile). It is the only extant species in the ancien ...

— from

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...

. All other American marsupials are members of the

Ameridelphia Ameridelphia is traditionally a superorder that includes all marsupials living in the Americas except for the Monito del monte (''Dromiciops''). It is now regarded as a paraphyletic group. Orders The orders within this group are listed below: ...

. Analysis of

retrotransposon Retrotransposons (also called Class I transposable elements or transposons via RNA intermediates) are a type of genetic component that copy and paste themselves into different genomic locations (transposon) by converting RNA back into DNA through ...

insertion sites in the

nuclear DNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. ...

of a variety of marsupials has shown that the South American

's lineage is the most

basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...

of the superorder. The Australian australidelphians form a clade, for which the name Euaustralidelphia ("true Australidelphia") has been proposed (the branching order within this group is yet to be determined). The study also showed that the most basal of all marsupial orders are the other two South American groups ( Didelphimorphia and Paucituberculata, with the former probably branching first). This indicates that Australidelphia arose in South America along with the other major divisions of extant marsupials, and likely reached Australia via Antarctica in a single

dispersal event Biological dispersal refers to both the movement of individuals ( animals, plants, fungi, bacteria, etc.) from their birth site to their breeding site ('natal dispersal'), as well as the movement from one breeding site to another ('breeding ...

after

Microbiotheria Microbiotheria is an australidelphian marsupial order that encompasses two families, Microbiotheriidae and Woodburnodontidae, and is represented by only one extant species, the monito del monte, and a number of extinct species known from foss ...

split off.

Phylogeny

Phylogeny of living Australidelphia based on the work of May-Collado, Kilpatrick & Agnarsson 2015 with extinct clades from Black et al. 2012

Taxonomy

The

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...

s within this group are listed below: * Genera †''

Djarthia ''Djarthia'' is an extinct monotypic genus of marsupial. It is the oldest marsupial found in Australia, discovered at the Murgon fossil site in south-eastern Queensland. ''D.'' ''murgonensis'' was described from material identified as Early ...

'' Godthelp, Wroe & Archer 1999 * Order † Yalkaparidontia Archer, Hand & Godthelp 1988 ** Family † Yalkaparidontidae Archer, Hand & Godthelp 1988 * ?Order

(1 species) ** ?Family

Microbiotheriidae Microbiotheriidae is a family of australidelphian marsupials represented by only one extant species, the monito del monte, and a number of extinct species known from fossils in South America, Western Antarctica, and northeastern Australia. Mi ...

* Order

Dasyuromorphia Dasyuromorphia (, meaning "hairy tail" in Greek) is an order comprising most of the Australian carnivorous marsupials, including quolls, dunnarts, the numbat, the Tasmanian devil, and the thylacine. In Australia, the exceptions include the ...

(71 species) ** Family †

Thylacinidae Thylacinidae is an extinct family of carnivorous, superficially dog-like marsupials from the order Dasyuromorphia. The only species to survive into modern times was the thylacine (''Thylacinus cynocephalus''), which became extinct in 1936. The ...

thylacine The thylacine ( , or , also ) (''Thylacinus cynocephalus'') is an extinct carnivorous marsupial that was native to the Australian mainland and the islands of Tasmania and New Guinea. The last known live animal was captured in 1930 in Tas ...

** Family

Dasyuridae The Dasyuridae are a family of marsupials native to Australia and New Guinea, including 71 extant species divided into 17 genera. Many are small and mouse-like or shrew-like, giving some of them the name marsupial mice or marsupial shrews, but ...

antechinus ''Antechinus'' (// ('ant-echinus')) is a genus of small dasyurid marsupial endemic to Australia. They resemble mice with the bristly fur of shrews. Names They are also sometimes called 'broad-footed marsupial mice', 'pouched mice', or 'Ante ...

es,

quoll Quolls (; genus ''Dasyurus'') are carnivorous marsupials native to Australia and New Guinea. They are primarily nocturnal and spend most of the day in a den. Of the six species of quoll, four are found in Australia and two in New Guinea. Anot ...

dunnart Dunnart is a common name for species of the genus ''Sminthopsis'', narrow-footed marsupials the size of a European mouse. They have a largely insectivorous diet. Taxonomy The genus name ''Sminthopsis'' was published by Oldfield Thomas in 1 ...

Tasmanian devil The Tasmanian devil (''Sarcophilus harrisii'') (palawa kani: purinina) is a carnivorous marsupial of the family Dasyuridae. Until recently, it was only found on the island state of Tasmania, but it has been reintroduced to New South Wales in ...

, and allies ** Family

Myrmecobiidae The numbat (''Myrmecobius fasciatus''), also known as the noombat or walpurti, is an insectivorous marsupial. It is diurnal and its diet consists almost exclusively of termites. The species was once widespread across southern Australia, but ...

numbat The numbat (''Myrmecobius fasciatus''), also known as the noombat or walpurti, is an insectivorous marsupial. It is diurnal and its diet consists almost exclusively of termites. The species was once widespread across southern Australia, but ...

* Order

Peramelemorphia The order Peramelemorphia includes the bandicoots and bilbies; it equates approximately to the mainstream of marsupial omnivores. All members of the order are endemic to the twin land masses of Australia-New Guinea and most have the characteris ...

(21 species) ** Family Thylacomyidae: bilbies ** Family †Chaeropodidae: pig-footed bandicoots ** Family

Peramelidae The marsupial family Peramelidae contains all of the extant bandicoots. They are found throughout Australia and New Guinea, with at least some species living in every available habitat, from rainforest to desert. Four fossil peramelids are descr ...

bandicoot Bandicoots are a group of more than 20 species of small to medium-sized, terrestrial, largely nocturnal marsupial omnivores in the order Peramelemorphia. They are endemic to the Australia–New Guinea region, including the Bismarck Archipelag ...

s and allies * Order

Notoryctemorphia Notoryctidae is a family of mammals, allying several extant and fossil species of Australia. The group appear to have diverged from other marsupials at an early stage and are highly specialised to foraging through loose sand; the unusual feature ...

(2 species) ** Family

Notoryctidae Notoryctidae is a family of mammals, allying several extant and fossil species of Australia. The group appear to have diverged from other marsupials at an early stage and are highly specialised to foraging through loose sand; the unusual feature ...

: marsupial moles * Order

Diprotodontia Diprotodontia (, from Greek "two forward teeth") is the largest extant order of marsupials, with about 155 species, including the kangaroos, wallabies, possums, koala, wombats, and many others. Extinct diprotodonts include the hippopotamus-sized ...

(117 species) ** Family

Phascolarctidae The Phascolarctidae (''φάσκωλος (phaskolos)'' - pouch or bag, ''ἄρκτος (arktos)'' - bear, from the Greek ''phascolos'' + ''arctos'' meaning pouched bear) is a family of marsupials of the order Diprotodontia, consisting of only on ...

koala The koala or, inaccurately, koala bear (''Phascolarctos cinereus''), is an arboreal herbivorous marsupial native to Australia. It is the only extant representative of the family Phascolarctidae and its closest living relatives are the ...

** Family Vombatidae:

wombat Wombats are short-legged, muscular quadrupedal marsupials that are native to Australia. They are about in length with small, stubby tails and weigh between . All three of the extant species are members of the family Vombatidae. They are ad ...

s ** Family

Phalangeridae The Phalangeridae are a family of mostly nocturnal marsupials native to Australia, New Guinea, and Eastern Indonesia, including the cuscuses, brushtail possums, and their close relatives. Considered a type of possum, most species are arbore ...

: brushtail possums and cuscuses ** Family

Burramyidae The pygmy possums are a family of small possums that together form the marsupial family Burramyidae. The five extant species of pygmy possum are grouped into two genera. Four of the species are endemic to Australia, with one species also co-occ ...

: pygmy possums ** Family Tarsipedidae:

honey possum The honey possum or noolbenger (''Tarsipes rostratus''), is a tiny species of marsupial that feeds on the nectar and pollen of a diverse range of flowering plants. Found in southwest Australia, it is an important pollinator for such plants as '' ...

** Family

Petauridae Petauridae is a family of possums containing 13 species: four species of trioks and striped possum (genus ''Dactylopsila''), eight species of wrist-winged glider (genus ''Petaurus''), and Leadbeater's possum (''Gymnobelideus leadbeateri''), whi ...

: striped possum,

Leadbeater's possum Leadbeater's possum (''Gymnobelideus leadbeateri'') is a critically endangered possum largely restricted to small pockets of alpine ash, mountain ash, and snow gum forests in the Central Highlands of Victoria, Australia, north-east of Melb ...

yellow-bellied glider The yellow-bellied glider (''Petaurus australis''), also known as the fluffy glider, is an arboreal and nocturnal gliding possum that lives in native eucalypt forests in eastern Australia, from northern Queensland south to Victoria. Habitat The ...

sugar glider The sugar glider (''Petaurus breviceps'') is a small, omnivorous, arboreal, and nocturnal gliding possum belonging to the marsupial infraclass. The common name refers to its predilection for sugary foods such as sap and nectar and its ab ...

, mahogany glider,

squirrel glider The squirrel glider (''Petaurus norfolcensis'') is a nocturnal gliding possum. The squirrel glider is one of the wrist-winged gliders of the genus ''Petaurus''. Habitat This species' home range extends from Bordertown near the South Australi ...

** Family

Pseudocheiridae Pseudocheiridae is a family of arboreal marsupials containing 17 extant species of ringtailed possums and close relatives. They are found in forested areas and shrublands throughout Australia and New Guinea. Characteristics Physically, they app ...

: ringtailed possums and allies ** Family

Potoridae Potoroidae is a family of marsupials, small Australian animals known as bettongs, potoroos, and rat-kangaroos. All are rabbit-sized, brown, jumping marsupials and resemble a large rodent or a very small wallaby. Taxonomy The potoroids are sm ...

: potoroos, rat kangaroos, bettongs ** Family Acrobatidae:

feathertail glider The feathertail glider (''Acrobates pygmaeus''), also known as the pygmy gliding possum, pygmy glider, pygmy phalanger, flying phalanger and flying mouse, is a species of marsupial native to eastern Australia. It is the world's smallest gliding ...

and feather-tailed possum ** Family

Hypsiprymnodontidae The Hypsiprymnodontidae are a family of macropods, one of two families containing animals commonly referred to as rat-kangaroos. The single known extant genus and species in this family, the musky rat-kangaroo, ''Hypsiprymnodon moschatus'', oc ...

musky rat-kangaroo The musky rat-kangaroo (''Hypsiprymnodon moschatus'') is a small marsupial found only in the rainforests of northeastern Australia. First described in the later 19th century, the only other species are known from fossil specimens. They are simil ...

** Family

Macropodidae Macropodidae is a family of marsupials that includes kangaroos, wallabies, tree-kangaroos, wallaroos, pademelons, quokkas, and several other groups. These genera are allied to the suborder Macropodiformes, containing other macropods, and ...

: kangaroos, wallabies, and allies ** Family †

Thylacoleonidae Thylacoleonidae is a family of extinct meat-eating marsupials from Australia, referred to as marsupial lions. The best known is ''Thylacoleo carnifex'', also called the marsupial lion. The clade ranged from the Late Oligocene to the Pleistocen ...

: marsupial lions ** Family †

Palorchestidae Palorchestidae is an extinct family of diprotodont marsupials whose members are sometimes referred to as marsupial tapirs due to their superficial similarity to true tapirs. Genera The family consists of the following four genera: * ''Propalor ...

: marsupial tapirs ** ** ** ** ** Family †

Diprotodontidae The Diprotodontidae are an extinct family of large herbivorous marsupials, endemic to Australia and New Guinea during the Oligocene through Pleistocene periods from 28.4 million to 40,000 years ago. The family consisted of large quadrupedal t ...

: giant wombats ** ** **

Footnotes

References

Australasian realm fauna Mammal superorders Extant Paleocene first appearances Marsupials of Australia Marsupials Metatheria Paleocene mammals of Australia Paleocene mammals of Oceania {{Marsupial-stub