''Enophrys diceraus'', the antlered sculpin, is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...

, the typical sculpins. This species occurs in the northern Pacific Ocean.

Taxonomy

''Enophrys diceraus'' was first formally described as ''Cottus diceraus'' in 1787 by the German

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...

with its type locality given as

Kamchatka The Kamchatka Peninsula (, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and western coastlines, respectively. Immediately offshore along the Pacific ...

. In 1829 the French zoologist

Georges Cuvier Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...

described ''Cottus claviger'', also from Kamchatka, and in 1839 the English zoologist

William Swainson William Swainson Fellow of the Linnean Society, FLS, Fellow of the Royal Society, FRS (8 October 1789 – 6 December 1855), was an English ornithologist, Malacology, malacologist, Conchology, conchologist, entomologist and artist. Life Swains ...

proposed a new

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

for Cuvier's species which he called ''

Enophrys ''Enophrys'' is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Enophrys'' was first proposed as a monospecific genus in 183 ...

''. Later Cuvier's ''C. claviger'' was found to be a

synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

of Pallas's ''C, diceraus'' but it is the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

of the genus ''Enophrys''. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''diceraus'' means "two-horned" a reference to the preopercular spines, one on each side of the head.

Description

''Enophrys diceraus'' has a long straight spine on the upper cheek, the preopercular spine which reaches past the operculum and has between 2 and 8 ancillary spines, or spinules. The

lacrimal bone The lacrimal bones are two small and fragile bones of the facial skeleton; they are roughly the size of the little fingernail and situated at the front part of the medial wall of the orbit. They each have two surfaces and four borders. Several bon ...

has a bifurcated flap which extends down over the

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

. There are 2

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

s which are supported by between 7 or 8 spines and between 12 and 15 soft rays while the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

has 10 or 13 soft rays. There are 2 well developed bony ridges on the top of the head. The large

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

scales are plate like and have 1 or 2 keels and a rough rera edge. This species reaches a maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

of .

Distribution and habitat

''Enophrys diceraus'' is found in the subarctic part of the Pacific Ocean from the northern

Sea of Japan The Sea of Japan is the marginal sea between the Japanese archipelago, Sakhalin, the Korean Peninsula, and the mainland of the Russian Far East. The Japanese archipelago separates the sea from the Pacific Ocean. Like the Mediterranean Sea, it ...

to Point Hope in

Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...

and in the

Chukchi Sea The Chukchi Sea (, ), sometimes referred to as the Chuuk Sea, Chukotsk Sea or the Sea of Chukotsk, is a marginal sea of the Arctic Ocean. It is bounded on the west by the Long Strait, off Wrangel Island, and in the east by Point Barrow, Alaska, ...

to Amchitka Island in the

Aleutians The Aleutian Islands ( ; ; , "land of the Aleuts"; possibly from the Chukchi ''aliat'', or "island")—also called the Aleut Islands, Aleutic Islands, or, before 1867, the Catherine Archipelago—are a chain of 14 main, larger volcanic island ...

. It is a

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...

species of coastal waters dound at depths between over substrates of sand, mud and rock.

Biology

''Enophrys diceraus'' eats benthic invertebrates, especially crustaceans and molluscs, although adults eat

brittlestar Brittlestar is the stage name of Stewart Reynolds, a Stratford, Ontario based comedian, writer, communications consultant, and online television show host. His campaign for KFC Canada was the world's most watched branded video on Facebook in th ...

s too, The adults move into shallow water to spawn, the juvenile settle at lengths between and they become sexually mature at . They form aggregations for feeding and spawning in the south of their range in

Peter the Great Bay The Peter the Great Gulf (Russian: Залив Петра Великого) is a gulf on the southern coast of Primorsky Krai, Russia, and the largest gulf of the Sea of Japan. The gulf extends for from the Russian–North Korean border, at the ...

and in central parts along the mainland Asian coast and off the southern part of

Sakhalin Sakhalin ( rus, Сахали́н, p=səxɐˈlʲin) is an island in Northeast Asia. Its north coast lies off the southeastern coast of Khabarovsk Krai in Russia, while its southern tip lies north of the Japanese island of Hokkaido. An islan ...

. These do not occur in the northern part of their range.

References

{{Taxonbar, from=Q2440775 Fish described in 1787 Taxa named by Peter Simon Pallas diceraus