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The buntings are a group of
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
birds forming the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Emberiza'', the only genus in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Emberizidae. The family contains 44 species. They are
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
-eating birds with stubby, conical bills.


Taxonomy

The family Emberizidae was formerly much larger and included the species now placed in the
Passerellidae New World sparrows are a group of mainly New World passerine birds, forming the family Passerellidae. They are seed-eating birds with conical bills, brown or gray in color, and many species have distinctive head patterns. Although they share th ...
(New World sparrows) and
Calcariidae Calcariidae is a small family (biology), family of passerine birds. It includes longspurs and snow buntings. There are six species in three genera worldwide, found mainly in North America and Eurasia. They are migratory and can live in a variety ...
(longspurs and snow buntings). Molecular phylogenetic studies found that the large family consisted of distinct clades that were better treated as separate families. The
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Emberiza'' is now the only genus placed in the family Emberizidae. The genus was introduced by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1758 in the
tenth edition Tenth may refer to: Numbers * 10th, the ordinal form of the number ten * One tenth, , or 0.1, a fraction, one part of a unit divided equally into ten parts. ** the SI prefix deci- ** tithe, a one-tenth part of something * 1/10 of any unit of me ...
of his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
''. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
was subsequently designated as the
yellowhammer The yellowhammer (''Emberiza citrinella'') is a passerine bird in the Emberiza, bunting family that is native to Palearctic, Eurasia and has been introduced species, introduced to New Zealand and Australia. Most European birds remain in the br ...
(''Emberiza citrinella''). The genus name ''Emberiza'' is from
Old German Old High German (OHG; ) is the earliest stage of the German language, conventionally identified as the period from around 500/750 to 1050. Rather than representing a single supra-regional form of German, Old High German encompasses the numerous ...
, a bunting. The origin of the English "bunting" is unknown. A 2008 genetic study found that three emberizid species that were placed in their own
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genera clustered within the ''Emberiza''. These were the
crested bunting The crested bunting (''Emberiza lathami'') is a species of bird in the family Emberizidae. It is sparrow sized with males having rufous wings and tail on a black body and females being less contrastingly coloured in dull brown. Both males and fem ...
(''Melophus lathami''), the
slaty bunting The slaty bunting (''Emberiza siemsseni'') is a species of bird in the family Emberizidae. Appearance An adult Slaty bunting measures 13 centimeters in length and weighs 20 grams. The plumages are brown and highly distinctive with unusual tail f ...
(''Latouchiornis siemsseni''), and the corn bunting (''Miliaria calandra''). All three species are now included in the genus ''Emberiza''. A large DNA-based study of the passerines published in 2019 found that the buntings are most closely related to the longspurs and snow buntings in the family
Calcariidae Calcariidae is a small family (biology), family of passerine birds. It includes longspurs and snow buntings. There are six species in three genera worldwide, found mainly in North America and Eurasia. They are migratory and can live in a variety ...
. Ornithologists
Edward Dickinson Edward Dickinson (January 1, 1803 – June 16, 1874) was an American politician from Massachusetts. He is also known as the father of the poet Emily Dickinson; their family home in Amherst, the Emily Dickinson Museum, is a museum dedicated to h ...
and
Leslie Christidis Leslie Christidis (born 30 May 1959), also simply known as Les Christidis, is an Australian ornithologist. His main research field is the evolution and systematics of birds. He has been director of Southern Cross University National Marine Scien ...
in the fourth edition of the ''
Howard and Moore Complete Checklist of the Birds of the World The ''Howard and Moore Complete Checklist of the Birds of the World'' is a book by Richard Howard and Alick Moore which presents a list of the bird species of the world. It was the first single-volume world bird list to include subspecies names, ...
'' chose to split up ''Emberiza'' and recognise the genera ''Fringillaria'', ''Melophus'', ''Granativora'', ''Emberiza'', and ''Schoeniclus''. Their example has not been followed by the online version of the ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'' nor by Frank Gill and David Donsker in the list of world birds that they maintain on behalf of the
International Ornithologists' Union The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", ...
. The
British Ornithologists' Union The British Ornithologists' Union (BOU) aims to encourage the study of birds (ornithology) around the world in order to understand their biology and aid their conservation. The BOU was founded in 1858 by Professor Alfred Newton, Henry Baker ...
has argued that splitting the genus provides little benefit and destabilizes the nomenclature. Species in the
New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...
genus ''
Passerina The genus ''Passerina'' is a group of birds in the cardinal family (Cardinalidae). Although not closely related to the buntings in the family Emberizidae, they are sometimes known as the North American buntings. The males show vivid colors i ...
'' include the word "bunting" in their common names, but are now classed in the family
Cardinalidae Cardinalidae (sometimes referred to as "cardinal-grosbeaks" or simply "cardinals") is a family of New World-Endemism, endemic passerine birds that consists of Cardinalis, cardinals, grosbeaks, and Passerina, buntings. It also includes several ot ...
. The family is divided into four major clades. The species in Clade I are mainly African while those in Clades II to IV are Palearctic: The
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below is based on a study published in 2021. The phylogenetic relationships of two African species, the brown-rumped bunting (''Emberiza affinis'') and Vincent's bunting (''Emberiza vincenti''), were not determined.


List of species

The genus contains 44 species. Extinct species have been described: * †
Long-legged bunting The long-legged bunting (''Emberiza alcoveri'') is an extinct flightless species of bunting. It was distinguishable by its long legs and short wings, and it inhabited Tenerife, one of the Canary Islands. It is one of the few flightless passerin ...
(''Emberiza alcoveri'') (
Late Quaternary The Holocene () is the current geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene together form the Qu ...
) * † '' Emberiza shaamarica'' (Late Pliocene of Central Asia) * † '' Emberiza polgardiensis'' (Upper Miocene of Hungary) * † '' Emberiza media'' (Pliocene of Hungary) * † '' Emberiza parva'' (Pliocene of Hungary) * †'' Emberiza gaspariskii'' (Pliocene of Hungary) * †'' Emberiza bartoki'' (Middle Miocene of Hungary) (Nomen dubium) ''Emberiza pannonica'' from the upper Miocene of Hungary is also referred to this genus, but was later found to be a member of
Muscicapidae The Old World flycatchers are a large family, the Muscicapidae, of small passerine birds restricted to the Old World (Europe, Africa and Asia), with the exception of several vagrants and two species, bluethroat (''Luscinia svecica'') and north ...
.


References


Further reading

''Buntings and Sparrows – A Guide to the Buntings and North American Sparrows'' by Urban Olsson and Jon Curson, illustrated by Clive Byers (1995)


External links


Bunting videos, photos and sounds
on the Internet Bird Collection * {{Taxonbar, from1=Q614481, from2=Q28486 Bird common names . Finches Taxa named by Carl Linnaeus Non-human celestial navigation