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The elk (: ''elk'' or ''elks''; ''Cervus canadensis'') or wapiti, is the second largest species within the
deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...
family,
Cervidae A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) a ...
, and one of the largest terrestrial
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s in its native range of North America and Central and
East Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
. The word "elk" originally referred to the European variety of the
moose The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...
, ''Alces alces'', but was transferred to ''Cervus canadensis'' by North American colonists. The name "wapiti" is derived from a
Shawnee The Shawnee ( ) are a Native American people of the Northeastern Woodlands. Their language, Shawnee, is an Algonquian language. Their precontact homeland was likely centered in southern Ohio. In the 17th century, they dispersed through Ohi ...
and
Cree The Cree, or nehinaw (, ), are a Indigenous peoples of the Americas, North American Indigenous people, numbering more than 350,000 in Canada, where they form one of the country's largest First Nations in Canada, First Nations. They live prim ...
word meaning "white rump", after the distinctive light fur around the tail region which the animals may fluff-up or raise to signal their agitation or distress to one another, when fleeing perceived threats, or among males courting females and
sparring Sparring is a form of training common to many combat sports. It can encompass a range of activities and techniques such as punching, kicking, grappling, throwing, wrestling or submission work dependent on style. Although the precise form varies, ...
for dominance. A similar trait is seen in other
artiodactyl Artiodactyls are placental mammals belonging to the order Artiodactyla ( , ). Typically, they are ungulates which bear weight equally on two (an even number) of their five toes (the third and fourth, often in the form of a hoof). The other t ...
species, like the
bighorn sheep The bighorn sheep (''Ovis canadensis'') is a species of Ovis, sheep native to North America. It is named for its large Horn (anatomy), horns. A pair of horns may weigh up to ; the sheep typically weigh up to . Recent genetic testing indicates th ...
,
pronghorn The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American ante ...
and the
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...
, to varying degrees. Elk dwell in open forest and forest-edge habitats, grazing on grasses and sedges and browsing higher-growing plants, leaves, twigs and bark. Male elk have large, blood- and nerve-filled
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) Family (biology), family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally fo ...
s, which they routinely
shed A shed is typically a simple, single-storey (though some sheds may have two or more stories and or a loft) roofed structure, often used for storage, for hobby, hobbies, or as a workshop, and typically serving as outbuilding, such as in a bac ...
each year as the weather warms. Males also engage in ritualized mating behaviors during the
mating season Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and ch ...
, including posturing to attract females, antler-wrestling (sparring), and ''bugling'', a loud series of throaty whistles, bellows, screams, and other vocalizations that establish dominance over other males and aim to attract females. Elk were long believed to belong to a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the European
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or Hart (deer), hart, and a female is called a doe or hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Ir ...
(''Cervus elaphus''), but evidence from many
mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
genetic studies, beginning in 1998, shows that the two are distinct
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
. The elk's wider rump-patch and paler-hued antlers are key morphological differences that distinguish ''C. canadensis'' from ''C. elaphus''. Although it is currently only native to North America, Central,
East East is one of the four cardinal directions or points of the compass. It is the opposite direction from west and is the direction from which the Sun rises on the Earth. Etymology As in other languages, the word is formed from the fact that ea ...
and
North Asia North Asia or Northern Asia () is the northern region of Asia, which is defined in geography, geographical terms and consists of three federal districts of Russia: Ural Federal District, Ural, Siberian Federal District, Siberian, and the Far E ...
, elk once had a much wider distribution in the past; prehistoric populations were present across
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
and into
Western Europe Western Europe is the western region of Europe. The region's extent varies depending on context. The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the Western half of the ancient Mediterranean ...
during the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
, surviving into the early
Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
in Southern Sweden and the
Alps The Alps () are some of the highest and most extensive mountain ranges in Europe, stretching approximately across eight Alpine countries (from west to east): Monaco, France, Switzerland, Italy, Liechtenstein, Germany, Austria and Slovenia. ...
. The now-extinct North American
Merriam's elk The Merriam's elk (''Cervus canadensis merriami'') is an extinct subspecies of elk once found in the arid lands of the southwestern United States (in Arizona, New Mexico and Texas), as well as in Mexico. From the first New World arrival of Europ ...
subspecies (''Cervus canadensis merriami'') once ranged south into
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
. The wapiti has also successfully
adapted In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the p ...
to countries outside of its natural range where it has been introduced, including
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
and
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
; the animal's adaptability in these areas may, in fact, be so successful as to threaten the sensitive
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
ecosystems and species it encounters. As a member of the Artiodactyla order (and distant relative of the
Bovidae The Bovidae comprise the family (biology), biological family of cloven-hoofed, ruminant mammals that includes Bos, cattle, bison, Bubalina, buffalo, antelopes (including Caprinae, goat-antelopes), Ovis, sheep and Capra (genus), goats. A member o ...
), elk are susceptible to several
infectious disease An infection is the invasion of tissue (biology), tissues by pathogens, their multiplication, and the reaction of host (biology), host tissues to the infectious agent and the toxins they produce. An infectious disease, also known as a transmis ...
s which can be transmitted to or from domesticated
livestock Livestock are the Domestication, domesticated animals that are raised in an Agriculture, agricultural setting to provide labour and produce diversified products for consumption such as meat, Egg as food, eggs, milk, fur, leather, and wool. The t ...
. Efforts to eliminate infectious diseases from elk populations, primarily by
vaccination Vaccination is the administration of a vaccine to help the immune system develop immunity from a disease. Vaccines contain a microorganism or virus in a weakened, live or killed state, or proteins or toxins from the organism. In stimulating ...
, have had mixed success. Some cultures revere the elk as having spiritual significance. Antlers and velvet are used in
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...
s in parts of Asia; the production of ground antler and velvet supplements is also a thriving
naturopathic Naturopathy, or naturopathic medicine, is a form of alternative medicine. A wide array of practices branded as "natural", "non-invasive", or promoting "self-healing" are employed by its practitioners, who are known as naturopaths. Difficult ...
industry in several countries, including the United States,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
and
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
. The elk is hunted as a game species, and their meat is lean and higher in
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
than
beef Beef is the culinary name for meat from cattle (''Bos taurus''). Beef can be prepared in various ways; Cut of beef, cuts are often used for steak, which can be cooked to varying degrees of doneness, while trimmings are often Ground beef, grou ...
or
chicken The chicken (''Gallus gallus domesticus'') is a domesticated subspecies of the red junglefowl (''Gallus gallus''), originally native to Southeast Asia. It was first domesticated around 8,000 years ago and is now one of the most common and w ...
.


Naming and etymology

By the 17th century, ''Alces alces'' (moose, called "elk" in Europe) had long been
extirpated Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinctions mark a chan ...
from the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
, and the meaning of the word "elk" to English-speakers became rather vague, acquiring a meaning similar to "large deer". The name ''wapiti'' is from the
Shawnee The Shawnee ( ) are a Native American people of the Northeastern Woodlands. Their language, Shawnee, is an Algonquian language. Their precontact homeland was likely centered in southern Ohio. In the 17th century, they dispersed through Ohi ...
and
Cree The Cree, or nehinaw (, ), are a Indigenous peoples of the Americas, North American Indigenous people, numbering more than 350,000 in Canada, where they form one of the country's largest First Nations in Canada, First Nations. They live prim ...
word (in
Cree syllabics Cree syllabics are the versions of Canadian Aboriginal syllabics used to write Cree language, Cree dialects, including the original syllabics system created for Cree and Ojibwe language, Ojibwe. There are two main varieties of syllabics for Cre ...
: or ), meaning "white rump". There is a subspecies of wapiti in
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
called the
Altai wapiti The Altai wapiti, sometimes called the Altai elk, or simply the Altai Deer, is a subspecies of ''Cervus canadensis'' found in the forest hills of southern Siberia, northwestern Mongolia, and northern Xinjiang province of China China, offi ...
(''Cervus canadensis sibiricus''), also known as the Altai maral. According to the ''
Oxford English Dictionary The ''Oxford English Dictionary'' (''OED'') is the principal historical dictionary of the English language, published by Oxford University Press (OUP), a University of Oxford publishing house. The dictionary, which published its first editio ...
'', the etymology of the word "elk" is "of obscure history". In
Classical Antiquity Classical antiquity, also known as the classical era, classical period, classical age, or simply antiquity, is the period of cultural History of Europe, European history between the 8th century BC and the 5th century AD comprising the inter ...
, the European ''Alces alces'' was known as and , words probably borrowed from a
Germanic language The Germanic languages are a branch of the Indo-European language family spoken natively by a population of about 515 million people mainly in Europe, North America, Oceania, and Southern Africa. The most widely spoken Germanic language, ...
or another language of northern Europe. By the 8th century, during the
Early Middle Ages The Early Middle Ages (or early medieval period), sometimes controversially referred to as the Dark Ages (historiography), Dark Ages, is typically regarded by historians as lasting from the late 5th to the 10th century. They marked the start o ...
, the moose was known as derived from the
Proto-Germanic Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the linguistic reconstruction, reconstructed proto-language of the Germanic languages, Germanic branch of the Indo-European languages. Proto-Germanic eventually developed from ...
: ''*elho-'', ''*elhon-'' and possibly connected with the . Later, the species became known in
Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman Conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English pe ...
as ''elk'', ''elcke'', or ''elke'', appearing in the Latinized form ''alke'', with the spelling ''alce'' borrowed directly from . Noting that ''elk'' "is not the normal phonetic representative" of the Old English ''elch'', the ''Oxford English Dictionary'' derives ''elk'' from , itself from . The American ''Cervus canadensis'' was recognized as a relative of the red deer (''Cervus elaphus'') of Europe, and so ''Cervus canadensis'' were referred to as "red deer".
Richard Hakluyt Richard Hakluyt (; 1553 – 23 November 1616) was an English writer. He is known for promoting the British colonization of the Americas, English colonization of North America through his works, notably ''Divers Voyages Touching the Discov ...
refers to North America as a "lande ... full of many beastes, as redd dere" in his 1584 ''Discourse Concerning Western Planting''. Similarly, John Smith (explorer), John Smith's 1616 ''A Description of New England'' referred to red deer. Sir William Talbot, 3rd Baronet, Sir William Talbot's 1672 English translation of John Lederer's Latin ''Discoveries'' likewise called the species "red deer", but noted in parentheses that they were "for their unusual largeness improperly termed Elks by ignorant people". Both Thomas Jefferson's 1785 ''Notes on the State of Virginia'' and David Bailie Warden's 1816 ''Statistical, Political, and Historical Account of the United States'' used "red deer" to refer to ''Cervus canadensis''.


Taxonomy

Members of the genus ''Cervus'' (and hence early relatives or possible ancestors of the elk) first appear in the fossil record 25 million years ago, during the Oligocene in
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
, but do not appear in the North American fossil record until the early Miocene. The extinct Irish elk (''Megaloceros'') was not a member of the genus ''Cervus'' but rather the largest member of the wider deer family (Cervidae) known from the fossil record. Until recently, red deer and elk were considered to be one species, ''Cervus elaphus'', with over a dozen subspecies. But mitochondrial DNA studies conducted in 2004 on hundreds of samples from red deer and elk subspecies and other species of the ''Cervus'' deer family, strongly indicate that elk, or wapiti, should be a distinct species, namely ''Cervus canadensis''. DNA evidence validates that elk are more closely related to Thorold's deer (''C. albirostris'') and even sika deer (''C. nippon'') than they are to the red deer. Elk and red deer produce fertile offspring in captivity, and the two species have freely inter-bred in
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
's Fiordland National Park. The cross-bred animals have resulted in the disappearance of virtually all pure elk blood from the area. Key morphological differences that distinguish ''C. canadensis'' from ''C. elaphus'' are the former's wider rump patch and paler-hued antlers.


Subspecies

There are numerous
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of elk described, with six from North America and four from Asia, although some Taxonomy (biology), taxonomists consider them different ecotypes or Race (biology), races of the same species (adapted to local environments through minor changes in appearance and behavior). Populations vary in antler shape and size, body size, coloration and mating behavior. DNA investigations of the Eurasian subspecies revealed that Phenotype, phenotypic variation in antlers, mane and rump patch development are based on "climatic-related lifestyle factors". Of the six subspecies of elk known to have inhabited North America in historical times, four remain, including the Roosevelt elk, Roosevelt (''C. canadensis roosevelti''), Tule elk, Tule (''C. c. nannodes''), Manitoban elk, Manitoban (''C. c. manitobensis'') and Rocky Mountain elk (''C. c. nelsoni''). The eastern elk (''C. c. canadensis'') and
Merriam's elk The Merriam's elk (''Cervus canadensis merriami'') is an extinct subspecies of elk once found in the arid lands of the southwestern United States (in Arizona, New Mexico and Texas), as well as in Mexico. From the first New World arrival of Europ ...
(''C. c. merriami'') subspecies have been extinct for at least a century. Four subspecies described from the Asian continent include the
Altai wapiti The Altai wapiti, sometimes called the Altai elk, or simply the Altai Deer, is a subspecies of ''Cervus canadensis'' found in the forest hills of southern Siberia, northwestern Mongolia, and northern Xinjiang province of China China, offi ...
(''C. c. sibiricus'') and the Tianshan wapiti (''C. c. songaricus''). Two distinct subspecies found in China,
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
, the Korea, Korean Peninsula and Siberia are the Manchurian wapiti (''C. c. xanthopygus'') and the Alashan wapiti (''C. c. alashanicus''). The Manchurian subspecies is darker, and more reddish, in coloration than other populations. The Alashan wapiti of northern Central China is the smallest of all the subspecies, has the lightest coloration, and is one of the least-studied. Recent DNA analyses suggest that there are no more than three or four total subspecies of elk. All American forms, aside from possibly the Tule and the Roosevelt's elk, seem to belong to one subspecies—''Cervus c. canadensis''; even the Siberian elk (''C. c. sibiricus'') is, more or less, physically identical to the American forms, and thus may belong to this subspecies, too. However, the Manchurian wapiti (''C. c. xanthopygus'') is clearly distinct from the Siberian forms, but not distinguishable from the Alashan wapiti. Still, due to the insufficient genetic material that rejects monophyly of ''C. canadensis'', some researchers consider it premature to include the Manchurian wapiti as a true subspecies of wapiti, and that it likely needs to be elevated to its own species, ''C. xanthopygus''. The Chinese forms (the Sichuan deer, Kansu red deer, and Tibetan red deer) also belong to the wapiti, and were not distinguishable from each other by mitochondrial DNA studies. These Chinese subspecies are sometimes treated as a distinct species, namely the Central Asian red deer (''Cervus hanglu''), which also includes the Kashmir stag. * North American group ** Roosevelt's elk (''C. c. roosevelti'') ** Tule elk (''C. c. nannodes'') ** Manitoban elk (''C. c. manitobensis'') ** Rocky Mountain elk (''C. c. nelsoni'') ** Eastern elk (''C. c. canadensis''; extinct) **
Merriam's elk The Merriam's elk (''Cervus canadensis merriami'') is an extinct subspecies of elk once found in the arid lands of the southwestern United States (in Arizona, New Mexico and Texas), as well as in Mexico. From the first New World arrival of Europ ...
(''C. c. merriami''; extinct) * Asian/Eastern group **
Altai wapiti The Altai wapiti, sometimes called the Altai elk, or simply the Altai Deer, is a subspecies of ''Cervus canadensis'' found in the forest hills of southern Siberia, northwestern Mongolia, and northern Xinjiang province of China China, offi ...
(''C. c. sibiricus'') ** Tian Shan wapiti (''C. c. songaricus'') ** Manchurian wapiti (''C. c. xanthopygus'') ** Alashan wapiti (''C. c. alashanicus'') ** Tibetan red deer (''C. c. wallichii'') File:Audubon-eastern-elk.jpg, Artist's depiction of eastern elk File:The deer of all lands (1898) Altai wapiti.png, Illustration of
Altai wapiti The Altai wapiti, sometimes called the Altai elk, or simply the Altai Deer, is a subspecies of ''Cervus canadensis'' found in the forest hills of southern Siberia, northwestern Mongolia, and northern Xinjiang province of China China, offi ...
File:The deer of all lands (1898) Bedford's deer.png, Illustration of Manchurian wapiti File:The deer of all lands (1898) Hangul.png, Illustration of Kashmir stag


Characteristics

Elk have thick bodies with slender legs and short tails. They have a shoulder height of with a nose-to-tail length of . Males are larger and weigh while females weigh . The largest of the subspecies is the Roosevelt elk (''C. c. roosevelti''), found west of the Cascade Range in the U.S. states of California, Oregon and Washington (U.S. state), Washington, and in the Canadian province of British Columbia. Roosevelt elk have been introduced into Alaska, where the largest males are estimated to weigh up to . More typically, male Roosevelt elk weigh around , while females weigh .. Male tule elk weigh while females weigh . The whole weights of adult male Manitoban elk range from . Females have a mean weight of . The elk is the second largest extant species of deer, after the
moose The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...
. Antlers are made of bone, which can grow at a rate of per day. While actively growing, a soft layer of highly vascularized skin known as velvet (antler covering), velvet covers and protects them. This is shed in the summer when the antlers have fully developed. Bull elk typically have around six tines on each antler. The Siberian and North American elk carry the largest antlers while the Altai wapiti has the smallest. Roosevelt bull antlers can weigh . The formation and retention of antlers are testosterone-driven. In late winter and early spring, the testosterone level drops, which causes the antlers to shed. During the fall, elk grow a thicker coat of hair, which helps to insulate them during the winter. Both male and female North American elk grow thin neck manes; females of other subspecies may not. By early summer, the heavy winter coat has been shed. Elk are known to rub against trees and other objects to help remove hair from their bodies. All elk have small and clearly defined rump patches with short tails. They have different coloration based on the seasons and types of habitats, with gray or lighter coloration prevalent in the winter and a more reddish, darker coat in the summer. Subspecies living in arid climates tend to have lighter colored coats than do those living in forests. Most have lighter yellow-brown to orange-brown coats in contrast to dark brown hair on the head, neck, and legs during the summer. Forest-adapted Manchurian and Alaskan wapitis have red or reddish-brown coats with less contrast between the body coat and the rest of the body during the summer months. Calves are born spotted, as is common with many deer species, and lose them by the end of summer. Adult Manchurian wapiti may retain a few orange spots on the back of their summer coats until they are older. This characteristic has also been observed in the forest-adapted European red deer.


Behavior and ecology

Elk are among the most Sociality, gregarious deer species. During the summer group size can reach 400 individuals. For most of the year, adult males and females are segregated into different herds. Female herds are larger while bulls form small groups and may even travel alone. Young bulls may associate with older bulls or female groups. Male and female herds come together during the mating season, which may begin in late August. Males try to intimidate rivals by vocalizing and displaying with their antlers. If neither bull backs down, they engage in antler wrestling, sometimes sustaining serious injuries. Bulls have a loud, high-pitched, whistle-like vocalization known as ''bugling'', which advertise the male's fitness over great distances. Unusual for a vocalization produced by a large animal, buglings can reach a frequency of 4000 Hz. This is achieved by blowing air from the glottis through the nasal cavities. Elk can produce deeper pitched (150 Hz) sounds using the larynx. Cows produce an alarm bark to alert other members of the herd to danger, while calves will produce a high-pitched scream when attacked.


Reproduction and life cycle

Female elk have a short Estrous cycle, estrus cycle of only a day or two, and matings usually involve a dozen or more attempts. By the autumn of their second year, females can produce one and, very rarely, two offspring. Reproduction is most common when cows weigh at least . Dominant bulls follow groups of cows during the Rut (mammalian reproduction), rut from August into early winter. A bull will defend his harem of 20 cows or more from competing bulls and predators. Bulls also dig holes in the ground called wallows, in which Self-anointing#Ungulates, they urinate and roll their bodies. A male elk's urethra points upward so that urine is sprayed almost at a right angle to the penis. The urine soaks into their hair and gives them a distinct smell which attracts cows. A bull interacts with cows in his harem in two ways: herding and courtship. When a female wanders too far away from the harem's range, the male will rush ahead of her, block her path and aggressively rush her back to the harem. Herding behavior is accompanied by a stretched out and lowered neck and the antlers laid back. A bull may get violent and hit the cow with his antlers. During courtship, the bull is more peaceful and approaches her with his head and antlers raised. The male signals his intention to test the female for sexual receptivity by flicking his tongue. If not ready, a cow will lower her head and weave from side to side while opening and closing her mouth. The bull will stop in response in order not to scare her. Otherwise, the bull will copiously lick the female and then mount her. Younger, less dominant bulls, known as "spike bulls", because their antlers have not yet forked, will harass unguarded cows. These bulls are impatient and will not perform any courtship rituals and will continue to pursue a female even when she signals him to stop. As such, they are less reproductively successful, and a cow may stay close to the big bull to avoid harassment. Dominant bulls are intolerant of spike bulls and will chase them away from their harems. The gestation period is eight to nine months and the offspring weigh around . When the females are near to giving birth, they tend to isolate themselves from the main herd, and will remain isolated until the calf is large enough to escape predators. Calves are born spotted, as is common with many deer species, and they lose their spots by the end of summer. After two weeks, calves are able to join the herd, and are fully weaned at two months of age. Elk calves are as large as an adult
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...
by the time they are six months old. Elk will leave their natal (birth) ranges before they are three years old. Males disperse more often than females, as adult cows are more tolerant of female offspring from previous years. Elk live 20 years or more in captivity but average 10 to 13 years in the wild. In some subspecies that suffer less predation, they may live an average of 15 years in the wild.


Migration

As is true for many species of deer, especially those in mountainous regions, elk Animal migration, migrate into areas of higher altitude in the spring, following the retreating snows, and the opposite direction in the fall. Hunting pressure impacts migration and movement. During the winter, they favor wooded areas for the greater availability of food to eat. Elk do not appear to benefit from thermal cover. The Greater Yellowstone Ecosystem elk herds comprise as many as 40,000 individuals. During the spring and fall, they take part in the longest elk migration in the continental U.S., traveling as much as between summer and winter ranges. The Teton herd consists of between 9,000 and 13,000 elk and they spend winters on the National Elk Refuge, having migrated south from the southern portions of Yellowstone National Park and west from the Shoshone National Forest, Shoshone and Bridger–Teton National Forests.


Diet

Elk are ruminants and therefore have four-chambered stomachs. Unlike white-tailed deer and moose, which are chiefly browsers, elk are similar to cattle in that they are primarily Grazing, grazers. But like other deer, they also Browsing (herbivory), browse. Elk have a tendency to do most of their feeding in the mornings and evenings, seeking sheltered areas in between feedings to digest. Their diets vary somewhat depending on the season, with native grasses being a year-round supplement, tree bark (e.g. Cedar (plant), cedar, wintergreen, eastern hemlock, sumac, jack pine, red maple, Rhus typhina, staghorn, and basswood) being consumed in winter, and sedges, forbs, and tree sprouts during the summer. Favorites of the elk include dandelions, Aster (genus), aster, hawkweed, Viola (plant), violets, clover, and the occasional mushroom. Elk consume an average of of vegetation daily. Particularly fond of aspen sprouts which rise in the spring, elk have had some impact on aspen groves which have been declining in some regions where elk exist. Range and wildlife managers conduct surveys of elk pellet groups to monitor populations and resource use. Research in the Greater Yellowstone Ecosystem has found that supplemental feeding of concentrated alfalfa pellets leads to significant alterations in the elks' microbiome. The elk gut microbiome is typically characterized by a diverse community of bacteria specialized in breaking down complex plant fibers and cellulose, whereas the supplementally fed gut microbiome may have less fiber-digesting bacteria. Therefore, transitioning from natural foraging to concentrated alfalfa pellets can cause changes in the gut microbiome that might affect the elk's ability to efficiently digest their natural diet or could potentially lead to imbalances that affect overall health.


Predators and defensive tactics

Predators of elk include wolf, wolves, coyotes, Brown bear, brown and American black bear, black Asian black bear, bears, cougars, and Siberian tigers. Coyote packs mostly prey on elk calves, though they can sometimes take a winter- or disease-weakened adult. In the Greater Yellowstone Ecosystem, which includes Yellowstone National Park, bears are the most significant predators of calves while healthy bulls have never been recorded to be killed by bears and such encounters can be fatal for bears. The killing of cows in their prime is more likely to affect population growth than the killing of bulls or calves. Elk may avoid predation by switching from grazing to browsing. Grazing puts an elk in the compromising situation of being in an open area with its head down, leaving it unable to see what is going on in the surrounding area. Living in groups also lessens the risk of an individual falling to predation. Large bull elk are less vulnerable and can afford to wander alone, while cows stay in larger groups for protection for their calves. Bulls are more vulnerable to predation by wolves in late winter, after they have been weakened by months of chasing females and fighting. Males that have recently lost their antlers are more likely to be preyed upon.


Parasites and disease

At least 53 species of protist and animal parasites have been identified in elk. Most of these parasites seldom lead to significant mortality among wild or captive elk. ''Parelaphostrongylus tenuis'' (brainworm or meningeal worm) is a parasitic nematode known to affect the spinal cord and brain tissue of elk and other species, leading to death. The definitive host is the white-tailed deer, in which it normally has no ill effects. Snails and slugs, the intermediate hosts, can be inadvertently consumed by elk during grazing. The liver fluke ''Fascioloides magna'' and the nematode ''Dictyocaulus viviparus'' are also commonly found parasites that can be fatal to elk. Chronic wasting disease, transmitted by a misfolded protein known as a prion, affects the brain tissue in elk, and has been detected throughout their range in North America. First documented in the late 1960s in mule deer, the disease has affected elk on game farms and in the wild in a number of regions. Elk that have contracted the disease begin to show weight loss, changes in behavior, increased watering needs, excessive salivation and urinating and difficulty swallowing, and at an advanced stage, the disease leads to death. No risks to humans have been documented, nor has the disease been demonstrated to pose a threat to domesticated cattle. In 2002, South Korea banned the importation of elk antler velvet due to concerns about chronic wasting disease. The Gram-negative bacteria, Gram-negative bacterial disease brucellosis occasionally affects elk in the Greater Yellowstone Ecosystem, the only place in the U.S. where the disease is still known to exist, though this can extend out to the Bighorn Mountains. In domesticated cattle, brucellosis causes infertility, abortions, and reduced milk production. It is transmitted to humans as undulant fever, producing influenza-like symptoms that may last for years. Though bison are more likely to transmit the disease to other animals, elk inadvertently transmitted brucellosis to horses in Wyoming and cattle in Idaho. Researchers are attempting to eradicate the disease through vaccinations and herd-management measures, which are expected to be successful. Nevertheless, research has been ongoing since 2002, and a successful vaccine has yet to be developed . A recent Autopsy, necropsy study of captive elk in Pennsylvania attributed the cause of death in 33 of 65 cases to either gastrointestinal parasites (21 cases, primarily ''Eimeria'' sp. and ''Strongylida, Ostertagia'' sp.) or bacterial infections (12 cases, mostly pneumonia). Elk hoof disease was first noticed in the state of Washington in the late 1990s in the Cowlitz River basin, with sporadic reports of deformed hooves. Since then, the disease has spread rapidly with increased sightings throughout southwest Washington and into Oregon. The disease is characterised by deformed, broken, or missing hooves and leads to severe lameness in elk. The primary cause is not known, but it is associated with Treponema, treponeme bacteria, which are known to cause digital dermatitis in commercial livestock. The mode of transmission is also not known, but it appears to be highly contagious among elk. Studies are being undertaken by government departments to determine how to halt or eliminate the disease.


Distribution and status

The elk ranges from central Asia through to Siberia and east Asia and in North America. They can be found in open deciduous woodlands, boreal forests, upland moors, mountainous areas and grasslands. The International Union for Conservation of Nature and Natural Resources (IUCN) list the species as least-concern species. The habitat of Siberian elk in Asia is similar to that of the Rocky Mountain subspecies in North America. During the Late Pleistocene their range was much more extensive, being distributed across Eurasia, with remains being found as far west as France. These populations are most closely related to modern Asian populations of the elk. Their range collapsed at the start of the Holocene, possibly because they were specialized to cold periglacial tundra-steppe habitat. When this environment was replaced largely by closed forest the red deer might have outcompeted the elk. Relict (biology), Relictual populations survived into the early Holocene (until around 3000 years ago) in southern Sweden and the Alps, where the environment remained favorable. Elk were also present in the early Holocene of central Alaska, where it was the main hunted animal along with bison, and survived in Yukon until around 1400 BP (550 AD).


Introductions and reintroductions

As of 2014, population figures for all North American elk subspecies were around one million. Prior to the European colonization of North America, there were an estimated 10 million on the continent. There are many past and ongoing examples of reintroduction into areas of the US. Elk were reintroduced in Michigan in 1918 after going extinct there in 1875. The Rocky Mountain elk subspecies was reintroduced by hunter-conservation organizations into the Appalachian region of the U.S. where the now extinct eastern elk once lived. They were reintroduced to Pennsylvania beginning in 1913 and throughout the mid-20th Century, and now remain at a stable population of approximately 1,400 individuals. Since the late 1990s, they were reintroduced and recolonized in the U.S. states, states of Wisconsin, Kentucky, North Carolina, Tennessee, Georgia (U.S. state), Georgia, Virginia and West Virginia. In the state of Kentucky, the elk population in 2022 had increased to over 15,000 animals. In 2016, a male elk, likely from the Great Smoky Mountains, Smoky Mountains population, was sighted in South Carolina for the first time in nearly 300 years. Once locally extinct, dispersing elk are now regularly spotted in Iowa, although a wild population has not yet established. Since 2015, elk have also been reintroduced in a number of other states, including Missouri, and introduced to the islands of Etolin Island, Etolin and Afognak in Alaska. Reintroduction of the elk into Ontario began in the early 20th century and is ongoing with limited success. Elk and red deer were introduced to Argentina in the early 20th century. There they are now considered an invasive species, encroaching on Argentinian ecosystems where they compete for food with the indigenous South Andean deer, Chilean huemul and other herbivores. This negative impact on native animal species has led the IUCN to identify the elk as one of the world's 100 worst invaders. The introduction of deer to New Zealand began in the middle of the 19th century, and current populations are primarily European red deer, with only 15 percent being elk. In 1905 18 American wapiti were released in George Sound in the Fiordland National Park. In 1949 the New Zealand American Fiordland Expedition was undertaken to study the descendants of this release. There is significant hybridization of elk with red deer. These deer have had an adverse impact on forest regeneration of some plant species, as they consume more palatable species, which are replaced with those that are less favored by the elk. The long-term impact will be an alteration of the types of plants and trees found, and in other animal and plant species dependent upon them. As in Chile and Argentina, the IUCN has declared that red deer and elk populations in New Zealand are an invasive species.


U.S. states by estimated elk population


Cultural references

Elk have played an important role in the cultural history of a number of peoples. Neolithic petroglyphs from Asia depict antler-less female elk, which have been interpreted as symbolizing life and sustenance. They were also frequently overlaid with boats and associated with rivers, suggesting they also represented paths to the underworld. Petroglyphs of elk were carved into cliffs by the Ancestral Puebloans of the southwestern U.S. hundreds of years ago. The elk was of particular importance to the Lakota people, Lakota and played a spiritual role in their society. The male elk was admired for its ability to attract mates, and Lakota men will play a courting flute imitating a bugling elk to attract women. Men used elks' antlers as love charms and wore clothes decorated with elk images. The Rocky Mountain elk is the List of U.S. state mammals, official state animal of Utah. An image of an elk and a moose appear on the state Seal of Michigan, seal and Flag of Michigan, flag of Michigan. The Benevolent and Protective Order of Elks (B.P.O.E.) chose the elk as its namesake because a number of its attributes seemed appropriate for cultivation by members of the fraternity. Jewel-encrusted, gold-mounted elk teeth are prized possessions of many members of the B.P.O.E.


Commercial uses

Although the 2006 National Survey from the U.S. Fish and Wildlife Service does not provide breakdown figures for each game species, hunting of wild elk is most likely the primary economic impact. While elk are not generally harvested for meat production on a large scale, some restaurants offer the meat as a specialty item and it is also available in some grocery stores. The meat is higher in
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
and lower in fat and cholesterol than beef, pork, and chicken. Elk meat is a good source of iron, phosphorus and zinc. A male elk can produce of Velvet antler, antler velvet annually and on ranches in the United States, Canada and New Zealand, it is collected and sold to markets in
East Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
, where it is used in medicine. Some cultures consider antler velvet to be an aphrodisiac. However, consuming velvet from elk in North America may be risky since velvet from animals infected with chronic wasting disease may contain prions that could result in a human getting variant Creutzfeldt–Jakob disease. Antlers are also used in artwork, furniture and other novelty items. All Asian subspecies, along with other deer, have been raised for their antlers in central and eastern Asia by Han Chinese, Turkic peoples, Tungusic peoples, Mongols, Mongolians, and Koreans. Elk farming, Elk farms are relatively common in North America and New Zealand. Native Americans have used elk hides for tepee covering, clothing and footwear. Since 1967, the Boy Scouts of America have assisted employees at the National Elk Refuge in Wyoming by collecting the antlers which are shed each winter. They are then auctioned, with 80% of the proceeds returned to the refuge. In 2010, of antlers were auctioned, which brought in over United States dollar, $46,000.


References


External links


Arizona Elk
Arizona Game and Fish Department
Rocky Mountain Elk Foundation

Smithsonian Institution - North American Mammals: ''Cervus'' (''elaphus'') ''canadensis''
{{Taxonbar, from=Q180404 Red deer Extant Pleistocene first appearances Fauna of Central Asia Fauna of East Asia Fauna of Siberia Herbivorous mammals Fauna of the Holarctic realm Mammals described in 1777 Mammals of Canada Mammals of the United States Mammals of East Asia Taxa named by Johann Christian Polycarp Erxleben National symbols of Djibouti