Eldon's Galaxias
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Eldon's galaxias (''Galaxias eldoni'') is an endangered
galaxiid The Galaxiidae are a family of mostly small freshwater fish in the Southern Hemisphere. The majority live in Southern Australia or New Zealand, but some are found in South Africa, southern South America, Lord Howe Island, New Caledonia, and the ...
fish endemic to
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
. One of several fishes in the genus ''
Galaxias ''Galaxias'' is a genus of small freshwater fish in the family Galaxiidae, and are frequently referred to as the galaxiids. These highly adaptable fish are typically found at temperate latitudes across the Southern Hemisphere. Galaxiids are s ...
'' found in
Central Otago Central Otago is an area located in the inland part of the Otago region in the South Island of New Zealand. The motto for the area is "A World of Difference". The area is dominated by mountain ranges and the upper reaches of the Clutha River ...
, it has a very small home range and is at risk of being driven to extinction by
trout Trout (: trout) is a generic common name for numerous species of carnivorous freshwater ray-finned fishes belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of which are members of the subfamily Salmoninae in the ...
introduced for recreational fishing.


Taxonomy

Eldon's galaxias was described in 1997 by New Zealand freshwater ichthyologist
Bob McDowall Robert Montgomery McDowall (15 September 1939 – 20 February 2011) was one of New Zealand's most prominent freshwater ichthyologists. Biography McDowall was born on 15 September 1939, the son of dairy scientist Frederick Henry McDowall and en ...
from specimens collected in 1995 and 1996. It is part of the '' Galaxias vulgaris'' species complex, and differs from '' Galaxias pullus'', another new species McDowall had found in the same area genetically and by its colour pattern: irregular stripes that continue across its back. The name recognises the ichthyologist G. A. (Tony) Eldon, who helped collect specimens and had retired not long before McDowall described this species.


Description

''Galaxias eldoni'' is an elongated, deep-bodied ''Galaxias'', usually 80 mm in length (although very large individuals can reach 155 mm). Its snout shape is variable depending on whether it lives on a stream or wetland habitat. It has large pectoral and pelvic fins, and a dorsal fin that starts forward of the anal fin. It is orange-brown to olive-brown with very variable dark brown markings that can be speckles, spots, or irregular bands, and its skin is heavily dusted with gold. Sometimes it has a small black patch behind the gill cover ( operculum). Unlike all other ''Galaxias'' (except '' G. divergens)'', it has only 15
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
rays and no
pyloric caeca Starfish or sea stars are star-shaped echinoderms belonging to the class Asteroidea (). Common usage frequently finds these names being also applied to ophiuroids, which are correctly referred to as brittle stars or basket stars. Starfish ...
.


Habitat and distribution

This species lives only in eastern waterways of
Otago Otago (, ; ) is a regions of New Zealand, region of New Zealand located in the southern half of the South Island and administered by the Otago Regional Council. It has an area of approximately , making it the country's second largest local go ...
, in wetlands and small streams that flow into the Taieri, Tokomairiro and Waipori rivers and below
Lake Mahinerangi Lake Mahinerangi is a lake in Otago formed when a dam was built on the Waipori River for hydroelectricity, hydroelectric generation. The lake is to the west of Dunedin in New Zealand's South Island. Construction of the dam started in 1903 and si ...
. These streams tend to be shallow, fast flowing and stony, ranging from lowland forested areas to alpine. ''G. eldoni'' lives among the gravels or under the banks of the waterway. It is a relatively good climber, able to wriggle up wet rocky slopes, and is able to negotiate waterfalls up to 3 m high. It tends to be found upstream of waterfalls that block the movement of introduced
salmonid Salmonidae (, ) is a family of ray-finned fish, the only extant member of the suborder Salmonoidei, consisting of 11 extant genera and over 200 species collectively known as "salmonids" or "salmonoids". The family includes salmon (both Atlantic a ...
fishes such as
brown trout The brown trout (''Salmo trutta'') is a species of salmonid ray-finned fish and the most widely distributed species of the genus ''Salmo'', endemic to most of Europe, West Asia and parts of North Africa, and has been widely introduced globally ...
,
rainbow trout The rainbow trout (''Oncorhynchus mykiss'') is a species of trout native to cold-water tributary, tributaries of the Pacific Ocean in North America and Asia. The steelhead (sometimes called steelhead trout) is an Fish migration#Classification, ...
, and brook char. Its original distribution was likely much larger, but trout species have eradicated it in all but the headwaters which make up its current home range.


Feeding

''Galaxias eldoni'' feeds on small freshwater invertebrates such as
stoneflies Plecoptera is an order of insects commonly known as stoneflies. Some 3,500 species are described worldwide, with new species still being discovered. Stoneflies are found worldwide, except Antarctica. Stoneflies are believed to be one of the most ...
and
mayflies Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the order ...
. They will also feed on larger prey like freshwater crayfish/
kōura ''Paranephrops'' is a genus of freshwater crayfish found only in New Zealand. They are known by the English common names freshwater crayfish and koura, the latter from their Māori language, Māori name of ''kōura''. Species The two species a ...
, and food that is carried by the current.


Life cycle

Unlike the more well-known '
whitebait Whitebait is a collective term for the immature fry of fish, typically between long. Such young fish often travel together in schools along coasts, and move into estuaries and sometimes up rivers where they can be easily caught using fine-mes ...
' ''Galaxias'' species, ''G. eldoni'' is not
diadromous Fish migration is mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousa ...
; its entire life is spent in fresh water. It is long-lived, 15 years or longer; some fully-grown individuals have been tagged and then recaptured 10 years later. It is slow growing, reaching maturity at one (males) or two years (females). Females produce low numbers of large eggs during October. Spawning occurs at the stream edge in
riffle A riffle is a shallow landform in a flowing channel. Colloquially, it is a shallow place in a river where water flows quickly past rocks. However, in geology a riffle has specific characteristics. Topographic, sedimentary and hydraulic indica ...
s, and eggs are laid directly into the stony stream bottom. Larvae hatch 4–6 weeks later. The larvae are large and are able to swim effectively; they do not disperse far from where they were spawned.


Conservation

This species is rated Nationally Endangered by the
Department of Conservation Department may refer to: * Departmentalization, division of a larger organization into parts with specific responsibility Government and military *Department (administrative division), a geographical and administrative division within a country, ...
and critically endangered by the
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
, due to its small number of streams it occupies (an area of just 0.69 km2) and the threats of invasive species. Because ''G. eldoni'' lays very few eggs and has large, poorly dispersing fry, it is vulnerable to predation by introduced
trout Trout (: trout) is a generic common name for numerous species of carnivorous freshwater ray-finned fishes belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of which are members of the subfamily Salmoninae in the ...
. Since the species was described in 1997 several small populations have been driven extinct from conversion of land to forestry or farming. The building of irrigation canals has allowed the closely related species ''Galaxias pullus'' to invade its range and hybridise with it. Currently there are protected areas for this species in the Lammerlaw Range, but they are vulnerable to being invaded by trout, or trout being deliberately introduced.


References


External links

*Eldon's galaxias discussed on RNZ ''Critter of the Week''
4 August 2017
{{Taxonbar, from=Q5254084 Eldon's galaxias Endemic freshwater fish of New Zealand Fish of the South Island Taxa named by Bob McDowall Eldon's galaxias