Ekgmowechashala
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Sioux The Sioux or Oceti Sakowin ( ; Dakota/ Lakota: ) are groups of Native American tribes and First Nations people from the Great Plains of North America. The Sioux have two major linguistic divisions: the Dakota and Lakota peoples (translati ...
: "little cat man") is an extinct genus of
primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
belonging to
Adapiformes Adapiformes is a group of early primates. Adapiforms radiated throughout much of the northern continental mass (now Europe, Asia and North America), reaching as far south as northern Africa and tropical Asia. They existed from the Eocene to the ...
.


Description and significance

With a weight of approximately , around tall and resembling a
lemur Lemurs ( ; from Latin ) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea ( ), divided into 8 Family (biology), families and consisting of 15 genera and around 100 existing species. They are Endemism, ...
, ''Ekgmowechashala'' is the only known North American primate of its time; it lived during the late
Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...
and early
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
.


Classification

The classification of this form has long been problematic. It was variously classified as a member of the extinct family
Omomyidae Omomyidae is a group of early primates that radiated during the Eocene epoch between about (mya). Fossil omomyids are found in North America, Europe & Asia, making it one of two groups of Eocene primates with a geographic distribution spanning ...
(related to
tarsier Tarsiers ( ) are haplorhine primates of the family Tarsiidae, which is the lone extant family within the infraorder Tarsiiformes. Although the group was prehistorically more globally widespread, all of the existing species are restricted to M ...
s) and the equally extinct
Plagiomenidae Colugos (), flying lemurs, or cobegos (), are arboreal gliding euarchontogliran mammals that are native to Southeast Asia. Their closest evolutionary relatives are primates. There are just two living species of colugos: the Sunda flying lemur ...
(related to
colugo Colugos (), flying lemurs, or cobegos (), are arboreal gliding euarchontogliran mammals that are native to Southeast Asia. Their closest evolutionary relatives are primates. There are just two living species of colugos: the Sunda flying lemur ...
s), but has been recently reassigned to
Adapiformes Adapiformes is a group of early primates. Adapiforms radiated throughout much of the northern continental mass (now Europe, Asia and North America), reaching as far south as northern Africa and tropical Asia. They existed from the Eocene to the ...
, the extinct relatives of
lemur Lemurs ( ; from Latin ) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea ( ), divided into 8 Family (biology), families and consisting of 15 genera and around 100 existing species. They are Endemism, ...
s and other
strepsirrhine Strepsirrhini or Strepsirhini (; ) is a suborder of primates that includes the lemuriform primates, which consist of the lemurs of Madagascar, galagos ("bushbabies") and pottos from Africa, and the lorises from India and Southeast Asia. Colle ...
s. A
cladistic analysis Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
by Ni et al. (2016) reaffirmed the adapiform placement of ''Ekgmowechashala'' by recovering it as sister group to '' Bugtilemur'', '' Gatanthropus'', and ''
Muangthanhinius ''Muangthanhinius'' is a genus of adapiform primate that lived in Asia during the late Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleo ...
'' in Ekgmowechashalidae.


Paleobiology

The shape of its teeth, and their likeness to those of
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the North American, northern or common raccoon (also spelled racoon) to distinguish it from Procyonina, other species of raccoon, is a mammal native to North America. It is the largest ...
s, indicate that it ate soft fruit provided by the warm forests of the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
during the early Miocene.


Fossils

Fossil evidence of ''Ekgmowechashala'' was discovered on the
Pine Ridge Indian Reservation The Pine Ridge Indian Reservation (), also called Pine Ridge Agency, is an Oglala Lakota Indian reservation located in the U.S. state of South Dakota, with a small portion extending into Nebraska. Originally included within the territory of the ...
, an
Oglala Sioux The Oglala (pronounced , meaning 'to scatter one's own' in Lakota) are one of the seven subtribes of the Lakota people who, along with the Dakota, make up the Očhéthi Šakówiŋ (Seven Council Fires). A majority of the Oglala live on the P ...
Native American reservation in
South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state, state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Dakota people, Dakota Sioux ...
.
Molars The molars or molar teeth are large, flat tooth, teeth at the back of the mouth. They are more developed in mammal, mammals. They are used primarily to comminution, grind food during mastication, chewing. The name ''molar'' derives from Latin, '' ...
were found in 1981 in the basin of
John Day River The John Day River is a tributary of the Columbia River, approximately long, in northeastern Oregon in the United States. It is known as the Mah-Hah River by the Cayuse people. Undammed along its entire length, the river is the fourth longest ...
, and these are in the collection of the
Burke Museum of Natural History and Culture The Burke Museum of Natural History and Culture (commonly as Burke Museum) is a natural history museum on the campus of the University of Washington, in Seattle, Washington, United States. It is administered by the UW College of Arts and Scien ...
; in the summer of 1997 John Zancanella of the
Bureau of Land Management The Bureau of Land Management (BLM) is an agency within the United States Department of the Interior responsible for administering federal lands, U.S. federal lands. Headquartered in Washington, D.C., the BLM oversees more than of land, or one ...
found a lower molar in the
John Day Fossil Beds National Monument John Day Fossil Beds National Monument is a U.S. national monument in Wheeler and Grant counties in east-central Oregon. Located within the John Day River basin and managed by the National Park Service, the park is known for its well-preserve ...
. ''Ekgmowechashala philotau'', known from material in
Nebraska Nebraska ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Ka ...
and South Dakota, was thought to be the only species of this genus, but material from
Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...
has been recently described as a new species, ''E. zancanellai''. A tooth from the Toledo Bend Ranch Local Fauna of far eastern
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
has been assigned to this genus.


See also

*
List of fossil primates This is a list of fossil primates—extinct primates for which a fossil record exists. Primates are generally thought to have evolved from a small, unspecialized mammal, which probably fed on insects and fruits. However, the precise source of the p ...


References

{{Taxonbar, from=Q5350406, from2=Q24175560 Prehistoric primate genera Miocene primates Oligocene primates Miocene mammals of North America Oligocene mammals of North America Prehistoric mammals of North America Primates of North America Paleogene geology of Oregon Paleogene geology of South Dakota John Day Fossil Beds National Monument Oligocene genus first appearances Miocene genus extinctions Fossil taxa described in 1963 Adapiformes