''Ekembo nyanzae'', originally classed as a species of ''
Proconsul
A proconsul was an official of ancient Rome who acted on behalf of a consul. A proconsul was typically a former consul. The term is also used in recent history for officials with delegated authority.
In the Roman Republic, military command, or ' ...
'', is a species of
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
primate
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
first discovered by
Louis Leakey
Louis Seymour Bazett Leakey (7 August 1903 – 1 October 1972) was a Kenyan-British palaeoanthropologist and archaeologist whose work was important in demonstrating that humans evolved in Africa, particularly through discoveries made at Olduvai ...
on
Rusinga Island
Rusinga Island, with an elongated shape approximately 10 miles (16 km) from end to end and 3 miles (5 km) at its widest point, lies in the eastern part of Lake Victoria at the mouth of the Winam Gulf. Part of Kenya, it is linked to Mbita ...
in 1942, which he published in ''Nature'' in 1943. It is also known by the name ''Dryopithecus africanus''. A joint publication of
Wilfrid Le Gros Clark and
Louis Leakey
Louis Seymour Bazett Leakey (7 August 1903 – 1 October 1972) was a Kenyan-British palaeoanthropologist and archaeologist whose work was important in demonstrating that humans evolved in Africa, particularly through discoveries made at Olduvai ...
in 1951, "The Miocene Hominoidea of East Africa", first defines ''Proconsul nyanzae''. In 1965 Simons and
Pilbeam replaced ''Proconsul'' with ''Dryopithecus'', using the same species names.
In 1967, Louis defined ''
Kenyapithecus africanus'' on seven fossils from
Rusinga Island
Rusinga Island, with an elongated shape approximately 10 miles (16 km) from end to end and 3 miles (5 km) at its widest point, lies in the eastern part of Lake Victoria at the mouth of the Winam Gulf. Part of Kenya, it is linked to Mbita ...
. He saw it as an ancestor of ''wickeri'' and also of man, with a date of 20 mya in the middle
Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
. Another fossil found by the VanCouverings on Rusinga in 1967 seemed to confirm ''africanus''. In 1969 Simons and Pilbeam moved ''Kenyapithecus africanus'' into ''Dryopithecus nyanzae''. By 1978 the genus had recovered from the Dryopithecine event and was back to ''Proconsul''. In that year Andrews moved Clark & Leakey's 1951 ''Sivapithecus africanus'' into ''Proconsul nyanzae''. In 2015, it was moved into the new genus ''
Ekembo
''Ekembo'' is an early ape (hominoid) genus found in 17- to 20-million-year-old sediments from the Miocene epoch. Specimens have been found at sites around the ancient Kisingiri volcano in Kenya on Rusinga Island and Mfangano Island in Lake Vict ...
''.
A more recent discovery by Ward ''et al.'' in 1999
and reclassification splits ''Kenyapithecus africanus'' away again and lumps it with ''Equatorius africanus'', which would move it to the subfamily
Afropithecinae with ''
Afropithecus turkanensis''. As ''Ekembo'', ''
Kenyapithecus'' may not be in the same clade as apes and humans, but as the older ''
Equatorius
''Equatorius'' is an extinct genus of kenyapithecine primate found in central Kenya at the Tugen Hills. Thirty-eight large teeth belonging to this middle Miocene hominid in addition to a mandibular and partially complete skeleton dated 15.58 M ...
'', it may be.
Morphology
''Ekembo nyanzae'' had a
dental formula of 2:1:2:3 on both the upper and lower jaw. The upper premolars of ''E. nyanzae'' were large. This species had a relatively thick enamel on the molars. The
mandible
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movabl ...
of this
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
was relatively robust. ''E. nyanzae'' had an average body mass of about .
Range
''E. nyanzae'' lived on the
continent
A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions are commonly regarded as continents. Ordered from largest in area to smallest, these seven ...
of
Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
and the fossils were found in areas that suggest it lived in a dry, open
woodland
A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
environment
Environment most often refers to:
__NOTOC__
* Natural environment, all living and non-living things occurring naturally
* Biophysical environment, the physical and biological factors along with their chemical interactions that affect an organism or ...
.
See also
* ''
Chororapithecus
''Chororapithecus'' is an extinct great ape from the Afar region of Ethiopia roughly 8 million years ago during the Late Miocene, comprising one species, ''C. abyssinicus''. It is known from 9 isolated teeth discovered in a 2005–2007 survey of ...
''
* ''
Dryopithecus
''Dryopithecus'' is a genus of extinct great apes from the middle–late Miocene boundary of Europe 12.5 to 11.1 million years ago (mya). Since its discovery in 1856, the genus has been subject to taxonomic turmoil, with numerous new species b ...
''
* ''
Nakalipithecus
''Nakalipithecus nakayamai'' is an extinct species of great ape from Nakali, Kenya, from about 9.9–9.8 million years ago during the Late Miocene. It is known from a right jawbone with 3 molar tooth, molars and from 11 isolated teeth, and the ...
''
* ''
Pierolapithecus
''Pierolapithecus catalaunicus'' is an extinct species of primate which lived about 13 million years ago during the Miocene in what is now Hostalets de Pierola, Catalonia, Spain, giving it its scientific name. It is believed by some to be a comm ...
''
* ''
Samburupithecus
''Samburupithecus'' is an extinct primate that lived in Kenya during the middle to late Miocene. The one species in this genus, ''Samburupithecus kiptalami'', is known only from a maxilla fragment dated to discovered in 1982 and formally descri ...
''
References
External links
* The History Files
Hominid Chronology*http://members.tripod.com/cacajao/equatorius_africanus.html
Mikko's Phylogeny archive
Proconsul nyanzae classification in the Taxonomicon site
*
Partial skeleton of Proconsul nyanzae from Mfangano Island, Kenya abstract of article by CV Ward in ''American Journal of Physical Anthropology'', Volume 90, Issue 1, Pages 77 – 111, shown on Wiley Interscience site.
{{Taxonbar, from=Q2563949
Prehistoric apes
Miocene primates of Africa
Fossil taxa described in 1950