Ebrachosuchus
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''Ebrachosuchus'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of basal
phytosaur Phytosaurs (Φυτόσαυροι in Greek, meaning 'plant lizard') are an extinct group of large, mostly semiaquatic Late Triassic archosauriform or basal archosaurian reptiles. Phytosaurs belong to the order Phytosauria and are sometimes ref ...
known from the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
(late
Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227.3 ...
stage) of
Bavaria Bavaria, officially the Free State of Bavaria, is a States of Germany, state in the southeast of Germany. With an area of , it is the list of German states by area, largest German state by land area, comprising approximately 1/5 of the total l ...
, southern
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. It is known only from the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
BSPG 1931 X 501, a complete
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
missing both
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
s. It was collected at Ebrach Quarry, bed number 9 from the late Carnian-aged Blasensandstein Member of the Hassberge Formation. It was first named by Oskar Kuhn in
1936 Events January–February * January 20 – The Prince of Wales succeeds to the throne of the United Kingdom as King Edward VIII, following the death of his father, George V, at Sandringham House. * January 28 – Death and state funer ...
and the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Ebrachosuchus neukami''.Kuhn, O. (1936). Weitere Parasuchier und Labyrinthodonten aus dem Blasensandstein des mittleren Keuper von Ebrach. ''Palaeontographica, Abteilung A'', 83:61-98. Hunt and Lucas (1991) mistakenly referred to '' Francosuchus angustifrons'' as ''Ebrachosuchus angustifrons'', and considered it and the other two '' Francosuchus'' species, '' F. broilii'' and '' F. latus'', to be
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
s of ''E. neukami''. Furthermore, they reassigned the species to '' Paleorhinus'', and
synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
ized ''Ebrachosuchus'' with the former.Hunt, A. P., and Lucas, S. G. (1991)
The ''Paleorhinus'' biochron and the correlation of the non-marine Upper Triassic of Pangaea
''Palaeontology'' 34(2):487-501.
Subsequent researches accepted this referral. More recently, a phylogenetic analysis found ''E. neukami'' to be more closely related to Mystriosuchinae than to ''Paleorhinus'' and thus the genus ''Ebrachosuchus'' was re-validated, while ''F. angustifrons'' was reassigned as ''P. angustifrons'' as it shares unique synapomorphies with the type species of ''Paleorhinus'' (''P. bransoni'') and was found to be its
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
. ''F. broilii'' and ''F. latus'' were also removed from ''E. neukami'', as they were found to be
nomina dubia In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
.


References

Phytosauria Prehistoric reptile genera Late Triassic reptiles of Europe Triassic Germany Fossils of Germany Fossil taxa described in 1936 Taxa named by Oskar Kuhn {{triassic-reptile-stub