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The Ebenaceae are a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s belonging to
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Ericales The Ericales are a large and diverse order of flowering plants in the asterid group of the eudicots. Well-known and economically important members of this order include tea and ornamental camellias, persimmon, ebony, blueberry, cranberry, l ...
. The family includes
ebony Ebony is a dense black/brown hardwood, coming from several species in the genus '' Diospyros'', which also includes the persimmon tree. A few ''Diospyros'' species, such as macassar and mun ebony, are dense enough to sink in water. Ebony is fin ...
and
persimmon The persimmon () is the edible fruit of a number of species of trees in the genus '' Diospyros''. The most widely cultivated of these is the Chinese and Japanese kaki persimmon, ''Diospyros kaki''. In 2022, China produced 77% of the world's p ...
among about 768
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of trees and shrubs. It is distributed across the
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
and warmer
temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
regions of the world. It is most diverse in the
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s of
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...
, India, Thailand, tropical Africa and tropical America. Many species are valued for their wood, particularly ebony, for fruit, and as
ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...
s.


Biology

The fruits contain
tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and Precipitation (chemistry), precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' is widel ...
s, a
plant defense against herbivory Plant defense against herbivory or host-plant resistance is a range of adaptations Evolution, evolved by plants which improve their fitness (biology), survival and reproduction by reducing the impact of herbivores. Many plants produce secondary ...
, so they are often avoided by animals when unripe. The ripe fruits of many species are a food source for diverse animal taxa. The
foliage A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, f ...
is consumed by insects. The plants may have a strong scent. Some species have aromatic wood. They are important and conspicuous trees in many of their native
ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...
s, such as lowland
dry forests The tropical and subtropical dry broadleaf forest is a habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-round, and may receive ...
of the former
Maui Nui Maui Nui is a modern Geology, geologists' name given to a prehistoric Hawaii, Hawaiian island and the corresponding modern Biogeography, biogeographic region. Maui Nui is composed of four modern islands: Maui, Molokai, Molokaʻi, Lanai, Lānaʻi, ...
in
Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
,
Caspian Hyrcanian mixed forests The Hyrcanian forests (; ) are a zone of lush lowland and montane forests covering about near the shores of the Caspian Sea in Iran and Azerbaijan. The forest is named after the ancient region of Hyrcania. The World Wide Fund for Nature refers t ...
, Khathiar–Gir dry deciduous forests, Louisiade Archipelago rain forests,
Madagascar lowland forests The Madagascar lowland forests or Madagascar humid forests are a tropical and subtropical moist broadleaf forests, tropical moist broadleaf forest ecoregion found on the eastern coast of the island of Madagascar, home to a plant and animal mix th ...
, Narmada Valley dry deciduous forests, New Guinea mangroves, and
South Western Ghats montane rain forests The South Western Ghats montane rain forests is an ecoregion in South India, covering the southern portion of the Western Ghats in Karnataka, Kerala and Tamil Nadu at elevations from . Annual rainfall in this ecoregion exceeds . Setting The e ...
. Ebony is a dense black wood taken from several species in the genus ''
Diospyros ''Diospyros'' is a genus of over 700 species of deciduous and evergreen trees and shrubs. The majority are native to the tropics, with only a few species extending into temperate regions. Individual species valued for their hard, heavy, dark timb ...
'', including ''
Diospyros ebenum Ceylon ebony or ''Diospyros ebenum ( Kaluwara) (''Tamil: கருங்காலி Karungaali), is a species of tree in the genus Diospyros and the family Ebenaceae. The tree produces valuable black wood. Description This middle-height eve ...
'' (Ceylon ebony, Indian ebony), ''
Diospyros crassiflora ''Diospyros crassiflora'', commonly known as Gaboon ebony, African ebony, Cameroon ebony, Nigeria ebony, West African ebony, and Benin ebony is a species of lowland-rainforest tree in the family Ebenaceae that is endemic to Western Africa. It is ...
'' (West African ebony, Benin ebony), and ''
Diospyros celebica ''Diospyros celebica'' (commonly known as black ebony or Makassar ebony) is a species of flowering tree in the family Ebenaceae that is endemic to the island of Sulawesi in Indonesia. The common name Makassar ebony originates from the main sea ...
'' (Makassar ebony). '' Diospyros tesselaria'' (Mauritius ebony) was heavily exploited by the Dutch in the 17th century.


Description

The family includes
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s and
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s. The leaves are usually alternately arranged, but some species have opposite or whorled leaves. The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
is usually a cyme of flowers, sometimes a
raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...
or a
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
, and some plants produce solitary flowers. Most species are
dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...
. The flower has 3 to 8 petals, which are joined at the bases. There are usually several single or paired
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s, which are often attached to the inner wall of the corolla. Female flowers have up to 8 stigmas. The calyx is persistent. The fruits are
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
-like or capsular. Like the wood of some species, the roots and bark may be black in color.Duangjai, S., et al. (2006)
Generic delimitation and relationships in Ebenaceae sensu lato: evidence from six plastid DNA regions.
''American Journal of Botany'' 93(12), 1808-27.


Etymology

The family name Ebenaceae is based on the genus name ''Ebenus'', published by
Otto Kuntze Carl Ernst Otto Kuntze (23 June 1843 – 27 January 1907) was a German botanist. Biography Otto Kuntze was born in Leipzig. An apothecary in his early career, he published an essay entitled ''Pocket Fauna of Leipzig''. Between 1863 and 1866, he ...
in 1891. It is a later homonym of ''
Ebenus ''Ebenus'' is a genus of flowering plants in the family Fabaceae. It belongs to the subfamily Faboideae The Faboideae are a subfamily of the flowering plant family Fabaceae or Leguminosae. An acceptable alternative name for the subfamily is ...
'' L., a genus already named in the family
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
, and is thus ''
nomen illegitimum ''Nomen illegitimum'' (Latin for illegitimate name) is a technical term used mainly in botany. It is usually abbreviated as ''nom. illeg.'' Although the International Code of Nomenclature for algae, fungi, and plants uses Latin terms as qualif ...
''. The plant that Kuntze had named ''Ebenus'' was accordingly reassigned to the genus ''Maba'', which in turn has since been included in the genus ''Diospyros''. Because the name Ebenaceae had become well known, having been used in major botanical references such as Bentham and Hooker's ''
Genera Plantarum ''Genera Plantarum'' is a publication of Swedish naturalist Carl Linnaeus (1707–1778). The first edition was issued in Leiden, 1737. The fifth edition served as a complementary volume to ''Species Plantarum'' (1753). Article 13 of the Internat ...
'', Engler and Prantl's ''Natürlichen Pflanzenfamilien'', and Hutchinson's ''Families of Flowering Plants'', it was conserved and is therefore legitimate.


Genera

During the last century, seven genera have been included in the family at one time or another. One
phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
reduced the family to four genera:


References

{{Authority control Ericales families