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The eastern sand darter (''Ammocrypta pellucida'') is a species of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
, a darter from the subfamily
Etheostomatinae Etheosomatidae is a species rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The famil ...
, part of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains more than ...
, which also contains the
perch Perch is a common name for fish of the genus ''Perca'', freshwater gamefish belonging to the family Percidae. The perch, of which three species occur in different geographical areas, lend their name to a large order of vertebrates: the Per ...
es, ruffes and pikeperches. The eastern sand darter is a relatively small fish, most plentiful in the
Mississippi Mississippi () is a state in the Southeastern region of the United States, bordered to the north by Tennessee; to the east by Alabama; to the south by the Gulf of Mexico; to the southwest by Louisiana; and to the northwest by Arkansas. Mis ...
and Ohio Rivers, as well as
Lake Champlain , native_name_lang = , image = Champlainmap.svg , caption = Lake Champlain-River Richelieu watershed , image_bathymetry = , caption_bathymetry = , location = New York/Vermont in the United States; and Quebec in Canada , coords = , type = , ...
and the
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five la ...
. It prefers sandy-bottomed streams and sandy shoals in the lakes. The eastern sand darter feeds on larvae of
black flies A black fly or blackfly (sometimes called a buffalo gnat, turkey gnat, or white socks) is any member of the family Simuliidae of the Culicomorpha infraorder. It is related to the Ceratopogonidae, Chironomidae, and Thaumaleidae. Over 2,200 speci ...
and other small insects in the water. They also feed on
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
in small portions since their small mouth size limits their gape ability. Its average body size is around 1.5-2 in long, and it breeds in the spring and summer in sandy-bottomed waters.


Description

The average lifespan for the eastern sand darter is about 2–3 years, but if the siltation and pollution continue to worsen, the lifespan will gradually decrease. ''A. pellucida'' reproduces in the sandy shoals at the bottom of the lakes and rivers it inhabits during the spring and summer when the water is at its warmest temperature. It reaches sexual maturity around age one and the males are able to mate once during the breeding season, while females mate twice, producing about 350 eggs with an average clutch size of 71 eggs. If siltation continues to worsen, many of the eggs will be smothered, resulting in reduced reproduction, further hindering the eastern sand darters' population count.


Distribution and habitat

The eastern sand darter can be found in many areas throughout the United States and into southern Canada. In addition to the Mississippi and Ohio Rivers, it can be found in great concentration in Lakes Champlain, Erie, Huron, Michigan and Ontario. They can also be found in the
St. Lawrence River The St. Lawrence River (french: Fleuve Saint-Laurent, ) is a large river in the middle latitudes of North America. Its headwaters begin flowing from Lake Ontario in a (roughly) northeasterly direction, into the Gulf of St. Lawrence, connecting ...
drainage in Canada along with the Lake Champlain drainage in Vermont south into New York, Pennsylvania, West Virginia, Kentucky, Illinois, and Indiana. Currently, the greatest concentration occurs in northern Alabama and eastern Tennessee. Distribution has slowly decreased over the last few decades, and two key factors could be the cause. The eastern sand darter requires clean sand substrates; hence, siltation is a major factor in their decline. Siltation decreases the quality of
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
for both
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
development and adult darters by decreasing the oxygen levels within the sand in which they bury themselves. Furthermore, silted habitats can cause adult darters to not fully burrow or decrease their time burrowed which then causes them to waste energy reserves. Channel or water flow alterations, nutrient enrichment, or any other habitat modification can completely change the amount and quality of the shifting sand bars which can then turn detrimental for the eastern sand darter. The increased construction of dams also has been a major factor contributing to the reduction of their habitats. Dams reduce river levels and flow, decrease oxygen levels in reservoir waters, and alter water temperatures, making it difficult for the eastern sand darter to reproduce at a comfortable temperature. Dam construction adds a whole new dimension to decreasing the reproduction and survival rate for ''A. pellucida''.


Feeding

This fish seeks its
prey Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
from the sand along the bottom of the stream or lake. Their prey items usually range in size up to 3/16 inches, but prey selection varies depending on the age of the darter. Juvenile eastern sand darters consume small crustaceans, while adults prefer midge larvae, blackfly larvae, mayflies, and caddisflies. It has several species of fish predators, including
rainbow trout The rainbow trout (''Oncorhynchus mykiss'') is a species of trout native to cold-water tributaries of the Pacific Ocean in Asia and North America. The steelhead (sometimes called "steelhead trout") is an anadromous (sea-run) form of the coast ...
, largemouth and smallmouth bass, and
rock bass The rock bass (''Ambloplites rupestris''), also known as the rock perch, goggle-eye, red eye, and black perch, is a freshwater fish native to east-central North America. This red eyed creature is a species of freshwater fish in the sunfish fam ...
. However, since it spends extensive time buried in the sand and it has a translucent body, the eastern sand darter usually tends to be protected from predators. Minnows are the eastern sand darters' biggest competitor for food, but little conflict arises between the two since the darter occupies the lowest depths of the rivers and lakes while the minnows occupy the upper water column.


Taxonomy

The Eastern sand darter was first formally described as ''Pleurolepsis pellucidus'' in 1863 by the American
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of October ...
Frederic Ward Putnam (1839-1915) with the
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
given as the Black River at
Elyria, Ohio Elyria ( ) is a city in the Greater Cleveland metropolitan statistical area and the county seat of Lorain County, Ohio, United States, located at the forks of the Black River in Northeast Ohio 23 miles southwest of Cleveland. As of the 2020 cen ...
. This species forms a clade with the
scaly sand darter The scaly sand darter (''Ammocrypta vivax'') is a species of freshwater ray-finned fish, a darter from the subfamily Etheostomatinae, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the ...
(''A. vivax'') and the Southern sand darter (''A. meridiana'').


References

{{Taxonbar, from=Q303369 Ammocrypta Freshwater fish of the United States Fish of the Eastern United States Taxa named by Frederic Ward Putnam Fish described in 1863 Taxonomy articles created by Polbot