Eastern Sand Darter
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The eastern sand darter (''Ammocrypta pellucida'') is a species of freshwater
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...
, a darter from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Etheostomatinae Etheostomatinae is a species-rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The family ...
, part of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains nearly 250 ...
, which also contains the
perch Perch is a common name for freshwater fish from the genus ''Perca'', which belongs to the family Percidae of the large order Perciformes. The name comes from , meaning the type species of this genus, the European perch (''P. fluviatilis'') ...
es, ruffes and pikeperches. The eastern sand darter is a relatively small fish, most plentiful in the
Mississippi Mississippi ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South regions of the United States. It borders Tennessee to the north, Alabama to the east, the Gulf of Mexico to the south, Louisiana to the s ...
and
Ohio River The Ohio River () is a river in the United States. It is located at the boundary of the Midwestern and Southern United States, flowing in a southwesterly direction from Pittsburgh, Pennsylvania, to its river mouth, mouth on the Mississippi Riv ...
s, as well as
Lake Champlain Lake Champlain ( ; , ) is a natural freshwater lake in North America. It mostly lies between the U.S. states of New York (state), New York and Vermont, but also extends north into the Canadian province of Quebec. The cities of Burlington, Ve ...
and the
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes spanning the Canada–United States border. The five lakes are Lake Superior, Superior, Lake Michigan, Michigan, Lake Huron, H ...
. It prefers sandy-bottomed streams and sandy shoals in the lakes. The eastern sand darter feeds on larvae of
black flies A black fly or blackfly (sometimes called a buffalo gnat, turkey gnat, or white socks) is any member of the family Simuliidae of the Culicomorpha infraorder. It is related to the Ceratopogonidae, Chironomidae, and Thaumaleidae. Over 2,200 speci ...
and other small insects in the water. They also feed on
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
in small portions since their small mouth size limits their gape ability. Its average body size is around 1.5-2 in long, and it breeds in the spring and summer in sandy-bottomed waters.


Description

The average lifespan for the eastern sand darter is about 2–3 years, but if the siltation and pollution continue to worsen, the lifespan will gradually decrease. ''A. pellucida'' reproduces in the sandy shoals at the bottom of the lakes and rivers it inhabits during the spring and summer when the water is at its warmest temperature. It reaches sexual maturity around age one and the males are able to mate once during the breeding season, while females mate twice, producing about 350 eggs with an average clutch size of 71 eggs. If siltation continues to worsen, many of the eggs will be smothered, resulting in reduced reproduction, further hindering the eastern sand darters' population count.


Distribution and habitat

The eastern sand darter can be found in many areas throughout the United States and into southern Canada. In addition to the Mississippi and Ohio Rivers, it can be found in great concentration in Lakes Champlain, Erie, Huron, Michigan and Ontario. They can also be found in the
St. Lawrence River The St. Lawrence River (, ) is a large international river in the middle latitudes of North America connecting the Great Lakes to the North Atlantic Ocean. Its waters flow in a northeasterly direction from Lake Ontario to the Gulf of St. Lawren ...
drainage in Canada along with the Lake Champlain drainage in Vermont south into New York, Pennsylvania, West Virginia, Kentucky, Illinois, and Indiana. Currently, the greatest concentration occurs in northern Alabama and eastern Tennessee. Distribution has slowly decreased over the last few decades, and two key factors could be the cause. The eastern sand darter requires clean sand substrates; hence, siltation is a major factor in their decline. Siltation decreases the quality of
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
for both
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
development and adult darters by decreasing the oxygen levels within the sand in which they bury themselves. Furthermore, silted habitats can cause adult darters to not fully burrow or decrease their time burrowed which then causes them to waste energy reserves. Channel or water flow alterations, nutrient enrichment, or any other habitat modification can completely change the amount and quality of the shifting sand bars which can then turn detrimental for the eastern sand darter. The increased construction of dams also has been a major factor contributing to the reduction of their habitats. Dams reduce river levels and flow, decrease oxygen levels in reservoir waters, and alter water temperatures, making it difficult for the eastern sand darter to reproduce at a comfortable temperature. Dam construction adds a whole new dimension to decreasing the reproduction and survival rate for ''A. pellucida''.


Feeding

This fish seeks its
prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...
from the sand along the bottom of the stream or lake. Their prey items usually range in size up to 3/16 inches, but prey selection varies depending on the age of the darter. Juvenile eastern sand darters consume small crustaceans, while adults prefer midge larvae, blackfly larvae, mayflies, and caddisflies. It has several species of fish predators, including
rainbow trout The rainbow trout (''Oncorhynchus mykiss'') is a species of trout native to cold-water tributary, tributaries of the Pacific Ocean in North America and Asia. The steelhead (sometimes called steelhead trout) is an Fish migration#Classification, ...
, largemouth and
smallmouth bass The smallmouth bass (''Micropterus dolomieu'') is a species of freshwater fish in the Centrarchidae, sunfish family (biology), family (Centrarchidae) of the order (biology), order Centrarchiformes. It is the type species of its genus ''Micropterus ...
, and
rock bass The rock bass (''Ambloplites rupestris''), also known as the rock perch, goggle-eye, red eye, and black perch, is a freshwater fish native to east-central North America. This red-eyed fish is a species of freshwater fish in the sunfish family (bi ...
. However, since it spends extensive time buried in the sand and it has a translucent body, the eastern sand darter usually tends to be protected from predators. Minnows are the eastern sand darters' biggest competitor for food, but little conflict arises between the two since the darter occupies the lowest depths of the rivers and lakes while the minnows occupy the upper water column.


Taxonomy

The Eastern sand darter was first formally described as ''Pleurolepsis pellucidus'' in 1863 by the
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, p ...
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...
Frederic Ward Putnam (1839–1915) with the type locality given as the Black River at
Elyria, Ohio Elyria ( ) is a city in Lorain County, Ohio, United States, and its county seat. It is located at the forks of the Black River (Ohio), Black River in Northeast Ohio, southwest of Cleveland. The population was 52,656 at the 2020 United States cens ...
. This species forms a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
with the scaly sand darter (''A. vivax'') and the Southern sand darter (''A. meridiana'').


References

{{Taxonbar, from=Q303369 Ammocrypta Freshwater fish of the United States Fish of the Eastern United States Taxa named by Frederic Ward Putnam Fish described in 1863 Taxonomy articles created by Polbot