''Dyskritodon'' ("tooth of unknown origin", from

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

δυσκρίτος, "''dyskritos''"Sigogneau-Russell, D. (1995) Two possibly aquatic triconodont mammals from the Early Cretaceous of Morocco. Acta Palaeontologica Polonica. 40(2), p.149-162.) is a genus of extinct

mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...

from the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...

, and possibly the

Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

. Of uncertain affinities, it is tentatively described as a

eutriconodont Eutriconodonta is an order (biology), order of early mammals. Eutriconodonts existed in Asia (including Insular India, pre-contact India), Africa, Europe, North America, North and South America during the Jurassic and the Cretaceous periods. The ...

Description

The

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

, ''D. amazighi'', is known from the

Ksar Metlili Formation The Ksar Metlili Formation is a geological formation in eastern High Atlas of Morocco, it is late Tithonian to Berriasian in age. It is approximately thick and primarily consists of mudstone and sandstone, with thin calcareous beds. One of these ...

in the

Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. They separate the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range, which stretches around through M ...

, dating to the

Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...

. It is known from several

molars The molars or molar teeth are large, flat tooth, teeth at the back of the mouth. They are more developed in mammal, mammals. They are used primarily to comminution, grind food during mastication, chewing. The name ''molar'' derives from Latin, '' ...

, about 1.85 mm long. These teeth are noted for being rather high and narrow crowned, bearing three main cusps that decrease in height posteriorly, as well as two minuscule mesial cusps. ''D. indicus'' is known from a single lower molar tooth from the

Kota Formation The Kota Formation is a geological Formation (geology), formation in India. The age of the Kota Formation is uncertain; it is commonly considered to date to the Early Jurassic, but some studies have suggested it may extend into the Middle Jurassi ...

, dating to the

Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). The Hettangian follows the Rhaetian (part of the Triass ...

Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 192.9 ±0.3 Ma and 184.2 ±0.3 Ma (million years ago). The Plie ...

. It is very similar to ''D. amazighi'', differing in being smaller (1.24 mm), having a shorter posterior root and some differences in the cusps.Prasad & Manhas (2002) Triconodont mammals from the Jurassic Kota Formation of India. Geodiversitas 24 (2), p.445-464 However, compared to ''D. amazighi'' molars, it is "drastically less complete".

Classification

Sigogneau-Russell referred the genus to

Eutriconodonta Eutriconodonta is an order (biology), order of early mammals. Eutriconodonts existed in Asia (including Insular India, pre-contact India), Africa, Europe, North America, North and South America during the Jurassic and the Cretaceous periods. The ...

on the basis of its basic

triconodont Eutriconodonta is an order of early mammals. Eutriconodonts existed in Asia (including pre-contact India), Africa, Europe, North and South America during the Jurassic and the Cretaceous periods. The order was named by Kermack ''et al.'' in 1973 ...

shape. However, she considered it aberrant enough to warrant a separate family within the clade. Further studies have similarly found it rather distinct from other eutriconodonts, and it is listed as an ''

incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...

'' taxon. It is nonetheless more "advanced" than morganucodontids and other basal

Mammaliformes Mammaliaformes ("mammalian forms") is a clade of synapsid tetrapods that includes the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts during the Late Triassic. It is defined as ...

. Due to the large temporal distance between both species, different environments and general incompleteness of the Indian material, there is also doubt as to whereas ''D. indicus'' is closely related to ''D. amazighi''. For now, its relationship to it is, too, provisory.

Biology

As it was found in relative abundance in marine deposits, ''D. amazighi'', like the contemporary ''

Ichthyoconodon ''Ichthyoconodon'' is an extinct genus of eutriconodont mammal from the Lower Cretaceous of Morocco. It is notable for having been found in a unique marine location, and the shape of its teeth suggests an unusual, potentially fish-eating ecologic ...

'', has been suggested to be an

aquatic Aquatic means relating to water; living in or near water or taking place in water; does not include groundwater, as "aquatic" implies an environment where plants and animals live. Aquatic(s) may also refer to: * Aquatic animal, either vertebrate ...

piscivore A piscivore () is a carnivorous animal that primarily eats fish. Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted repti ...

. Similarities between its molars and those of mammals such as

seals Seals may refer to: * Pinniped, a diverse group of semi-aquatic marine mammals, many of which are commonly called seals, particularly: ** Earless seal, or "true seal" ** Fur seal * Seal (emblem), a device to impress an emblem, used as a means of a ...

and

cetaceans Cetacea (; , ) is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively c ...

have corroborated this hypothesis. Posterior analysis have shown a lack of precise equivalency between eutriconodont molars and those of

theria Theria ( or ; ) is a scientific classification, subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the Placentalia, placental mammals) and the metatherians (including the marsupials) but excludes the egg-lay ...

n mammals, rending this assessment as a fish-eater cautious, but the high state of preservation of the animal's teeth indicates that it died in situ or nearby, in open waters. ''D. indicus'', by contrast, appears in a terrestrial environment.

Notes

References

{{Taxonbar, from=Q8771060 Eutriconodonta Cretaceous mammals of Africa Cretaceous Morocco Fossils of Morocco Jurassic mammals of Asia Jurassic India Fossils of India Fossil taxa described in 1995 Taxa named by Denise Sigogneau-Russell Prehistoric mammal genera