Drymocallis Fissa
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''Drymocallis fissa'', the bigflower cinquefoil, also known as the leafy cinquefoil, leafy drymocallis, or wood beauty, is a small plant also sometimes classified as ''Potentilla fissa''. It is a herbaceous plant with a thick taproot known for its moderately hairy leaves, redish leaf stems, and relatively large yellow flowers. It is native to foothills and lower mountains the Rocky Mountain region in the western United States.


Description

Individuals of this species have numerous stems that come from the top of a single rootstalk with branched persistent woody stems (
caudex A caudex (: caudices) of a plant is a stem, but the term is also used to mean a rootstock and particularly a basal stem structure from which new growth arises.pages 456 and 695 In the strict sense of the term, meaning a stem, "caudex" is most ...
). The leaves are
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...
with 7-13 slightly hairy leaflets. Leaflet edges are incised to small points. The stems are hairy and redish. Plant usually most often has a cyme with multiple flowers per stem that is taller than the leaves coming up from the plant's base. Plants usually grow between 15–35 cm tall, but occasionally will be as short as 12 cm or as tall as 45 cm.


Flowers and fruit

The flowers are large as the common name suggests, 1.9 to 2.5 cm across, with five petals, colored bright yellow to creamy yellow. The center of the flower is also yellow and the
anthers The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filamen ...
are long, helping to distinguish it from other species. In Colorado ''Drymocallis fissa'' blooms in late spring to the very beginning of summer. The dry seed heads follow in July.


Taxonomy

This species was first described as ''Potentilla fissa'' by the widely cited botanist
Thomas Nuttall Thomas Nuttall (5 January 1786 – 10 September 1859) was an English botanist and zoologist who lived and worked in America from 1808 until 1841. Nuttall was born in the village of Long Preston, near Settle in the West Riding of Yorkshire a ...
in 1840, placing it with the other species in ''
Potentilla ''Potentilla'' is a genus containing over 500 species of Annual plant, annual, Biennial plant, biennial and Perennial plant, perennial herbaceous plant, herbaceous flowering plants in the rose family (biology), family, Rosaceae. Potentillas m ...
''. In 1898 the botanist
Per Axel Rydberg Per Axel Rydberg (July 6, 1860 – July 25, 1931) was a Swedish-born, American botanist who was the first curator of the New York Botanical Garden Herbarium. Biography Per Axel Rydberg was born in Odh, Västergötland, Sweden and emigrated to ...
published a paper arguing for a new classification as ''Drymocallis fissa''. While Rydberg's classification is widely used, Nuttall's placement in ''Potentilla'' continues to be used in some books in the 21st century.


Names

The genus ''Drymocallis'' was named by Rydberg using botanical Greek, "drymos" meaning woods and "kallos" meaning beauty. This also gives many plants in the genus as "wood beauty" or "woodbeauty". The previous classification as in the genus ''Cinquefoil'' also has given rise to common names for this species, such as "bigflower cinquefoil" and "leafy cinquefoil".


Distribution

'' Drymocallis fissa'' can be found from northern New Mexico to Alberta, Canada, but is most commonly found in the Front Range mountains and lower foothills in Colorado and Medicine Bow Mountains of Wyoming. It has also been reliably observed in the
Black Hills The Black Hills is an isolated mountain range rising from the Great Plains of North America in western South Dakota and extending into Wyoming, United States. Black Elk Peak, which rises to , is the range's highest summit. The name of the range ...
of South Dakota. The USDA records observations in
San Miguel County, New Mexico San Miguel County () is a county in the U.S. state of New Mexico. As of the 2020 census, the population was 27,201. Its county seat is Las Vegas. San Miguel County comprises the Las Vegas Micropolitan Statistical Area, which is also include ...
;
Carbon County, Montana Carbon County is a county located in the U.S. state of Montana. As of the 2020 census, the population was 10,473. Its county seat is Red Lodge. Carbon County is part of the Billings, MT Metropolitan Statistical Area. History Carbon Count ...
;
Stillwater County, Montana Stillwater County is a county in the U.S. state of Montana. As of the 2020 census, the population was 8,963. Its county seat is Columbus. Geography According to the United States Census Bureau, the county has a total area of , of which is ...
; and
Custer County, Idaho Custer County is a rural mountain county in the center of the U.S. state of Idaho. As of the 2020 census, the population was 4,275, making it the fifth-least populous county in Idaho. The county seat is Challis. Established in 1881, the cou ...
as well as Oregon and Utah, but these are suspected of being misidentifications of other species.


Habitat and ecology

Bigflower cinquefoil is a perennial plant that often grows in mixed woodland habitats in the lower foothills, with sagebrush, on rocky slopes, and in open meadows at higher elevations. It is found from 1600–3000 meters in altitude. It prefers disturbed and rocky areas with more water, but not waterlogged soils. ''Drymocallis fissa'' was evaluated by
NatureServe NatureServe, Inc. is a non-profit organization based in Arlington County, Virginia, United States, US, that provides proprietary wildlife conservation-related data, tools, and services to private and government clients, partner organizations, and ...
as apparently secure (G4) in 1994. This means that at a global level they think it has fairly low risk of extinction or collapse due to an extensive range and/or many populations, but with possible cause for some concern. It was additionally rated as vulnerable (S3) in Wyoming.


References

{{Taxonbar, from1=Q17246156, from2=Q50346929 fissa Flora of Colorado Flora of South Dakota Flora of Wyoming Plants described in 1840