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Dothideomycetidae is a fungal subclass in the class
Dothideomycetes Dothideomycetes is the largest and most diverse class of ascomycete fungi. It comprises 11 orders 90 families, 1,300 genera and over 19,000 known species. Wijayawardene et al. in 2020 added more orders to the class. Traditionally, most of it ...
The cavities of the sexual structures do not have vertical cells ( paraphyses, pseudoparaphyses or paraphysoids) growing between the sac-like cells bearing the sexual spores ( asci).


Description

Members of the Dothideomycetidae form small to medium fruiting bodies ( ascomata) that develop either within the host tissue or substrate or, in some species, burst through to become visible at the surface; a minority are superficially seated from the outset. Each ascoma may comprise a single cavity (locule) or several interconnected chambers where the spore-bearing sacs ( asci) are housed. A minute pore (the ostiole) usually connects the cavity to the outside air, and its short lining filaments () can sometimes be seen under the microscope. The surrounding wall is often built of densely packed, brick-like fungal cells (), but the inner space lacks the long sterile threads () found in many other ascomycetes. The asci themselves display a wide range of outlines—from nearly spherical through egg-shaped and club-shaped to elongated cylinders. Inside, the
spore In biology, a spore is a unit of sexual reproduction, sexual (in fungi) or asexual reproduction that may be adapted for biological dispersal, dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores fo ...
s vary from colourless to dark brown and may be single-celled, divided by one or several cross walls ( septa), or partitioned in both directions to give a (brick-work) appearance. In addition to
sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote tha ...
, Dothideomycetidae exhibit asexual stages: they may produce
conidia A conidium ( ; : conidia), sometimes termed an asexual chlamydospore or chlamydoconidium (: chlamydoconidia), is an asexual, non- motile spore of a fungus. The word ''conidium'' comes from the Ancient Greek word for dust, ('). They are also ...
inside flask-like chambers embedded in tissue (coelomycetous anamorphs) or openly on thread-like hyphae ( hyphomycetous anamorphs).


Classification

According to ''2024 Outline of Fungi'', the Dothideomycetidae consists of the following orders: * Arthrocatenales * Aureoconidiellales * Capnodiales * Cladosporiales * Comminutisporales *
Dothideales Dothideales are an order (biology), order of bitunicate fungi consisting mainly of saprobic or plant parasitic species. Description Taxa in this order are characterized by the absence of a hamathecium (defined as hyphae or other tissues betwee ...
* Mycosphaerellales * Myriangiales * Neophaeothecales * Phaeothecales * Racodiales


Genera ''incertae cedis''

*'' Aenigmatomyces''


References

{{Taxonbar, from=Q146132 Dothideomycetes Fungus subclasses Lichen subclasses