The Dixidae (meniscus
midge
A midge is any small fly, including species in several family (biology), families of non-mosquito nematoceran Diptera. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid ...
s) are a
family
Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
aquatic nematocera
The Nematocera (the name meaning "thread-horns") are a suborder of elongated fly, flies with thin, segmented antenna (biology), antennae and mostly aquatic larvae. This group is paraphyletic and contains all flies except for species from suborder ...
n
flies
Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...
(Diptera). The
larva
A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e live in unpolluted, standing
fresh water
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salt (chemistry), salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include ...
s, just beneath the surface film, usually amongst marginal aquatic vegetation. They are found in all continents except Antarctica.
Description

Dixidae are small (body length not more than 5.0 mm) slender gnats with thin legs. Adults are black to yellowish-brown. The head is relatively broad. The antennae are thin and the flagellum has 14 segments. The proboscis is short and thick and the palpi are five-segmented. The thorax is slightly convex. The wing veins are without scales (with scales in the closely related family
Culicidae
Mosquitoes, the Culicidae, are a Family (biology), family of small Diptera, flies consisting of 3,600 species. The word ''mosquito'' (formed by ''Musca (fly), mosca'' and diminutive ''-ito'') is Spanish and Portuguese for ''little fly''. Mos ...
). The subcosta is fused with the costa at the level of the base of Rs or slightly proximal to this. The wing venation exhibits radial, medial, and cubital forks (R 4 branched, M 2 branched, Cu 2 branched). R 2+ 3 is strongly arched, the r–m crossvein is distinct, and the discal cell is absent. The anal vein of the wing is long. The genitalia of the male is inverted at 180° by torsion of segments 5–8.
Evolutionary history
The oldest known fossils of the group come from the
Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
of Asia, assigned to the extinct genera ''
Syndixa'' and ''
Eucorethrina,'' members of modern genera are not known until the
Eocene
The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
.
References
Further reading
*
Lindner, E 193 1. Dixinae (Culicidae) 3, 11–12, 1-43 In: Lindner, E. (Ed.). ''Die Fliegen der Paläarktischen Region'' Keys to Palaearctic species but now needs revision (in German).
*A. A. Shtakel'berg Family Dixidae in
Bei-Bienko, G. Ya, 1988 ''Keys to the insects of the European Part of the USSR'' Volume 5 (Diptera) Part 2 English edition.Keys to Palaearctic species but now needs revision .
External links
Delta Int Keyfamily description and image
BugGuideWing venationEOLimages
Diptera.infoimages
*
Dixiidae in Italian
Nematocera families
{{Culicoidea-stub