Dipteryx

''Dipteryx'' is a genus containing a number of species of large trees and possibly shrubs. It belongs to the "papilionoid" subfamily – Faboideae – of the family Fabaceae. This genus is native to South and Central America and the Caribbean. Formerly, the related genus '' Taralea'' was included in ''Dipteryx''.

Description

The largest members of ''Dipteryx'' are canopy-emergent trees of tropical rainforests. The tonka bean (''D. odorata'') is grown for its fragrant seeds. ''Baru'' (''D. alata'') is the only species which found in drier, seasonal areas, growing in the cerrado of Brazil; its fruit and seeds are used as food and fodder. Several species are used for timber, of which almendro (''D. oleifera'') wood is considered desirable, especially locally.

''Dipteryx'' can be distinguished from other members of the Dipterygeae by its compound leaves with asymmetric leaflets caused due to an eccentric primary vein, a drupaceous fruit, seeds with a leathery skin, a hilum in a lateral or subapical position and a rugose embryo with a conspicuous plumule.

Taxonomy

The number of recognised species of ''Dipteryx'' has changed over the years.

The genus was previously known as ''Coumarouna''. In 1934 Walter Adolpho Ducke split this genus into two, on the basis of the alternate leaflets, among other characters, of ''Dipteryx''. He used two older, conserved names published previously: ''Taralea'' and ''Dipteryx''. Although ''Taralea'' was accepted, some taxonomists did not recognise ''Dipteryx'' as the correct name for the genus until at least the mid-1940s.

In the most recent monograph on the genus, ''A Checklist of the Dipterygeae species'' by the Brazilian researcher Haroldo Cavalcante de Lima in 1989, he synonymised a number of species, accepting nine species in the genus. His taxonomy was accepted by ILDIS (2005) but not noticed or followed by US databases, i.e. in GRIN (2005), the entry on ''Dipteryx'' in the ''Contribución al conocimiento de las leguminosas Colombianas'' by C. Barbosa (1994), the IUCN (1998) based on ''World List of Threatened Trees'' by Oldfield ''et al''. (1998), or the ''Catalogue of the Flowering Plants and Gymnosperms of Peru'' (1993) which was built using the Tropicos database by the Missouri Botanical Garden. In 1999 the entry on ''Dipteryx'' in the ''Flora of the Venezuelan Guyana'' by de Lima was published.

The northernmost taxon '' Dipteryx panamensis'', notable as being the only species listed on CITES since 2003 and therefore subject to export controls, was synonymised with the neglected but older name ''D. oleifera'' by de Lima in 1989, but this move was only followed by ILDIS and one or two of articles on the species over the years, all other floras, databases and publications using the name ''D. panamensis''. In 2011, however, the ''Report of the Nomenclature Committee for Vascular Plants: 62'' recommended ''D. oleifera'' by treated as validly published, and de Lima's synonymy for this taxon has been accepted by many.

By 2010, in the ''Catálogo de Plantas e Fungos do Brasil'', de Lima had changed his mind and re-recognised two of Brazilian taxa he had earlier considered synonyms in 1989, although not all.

Species

12 species are accepted:
*'' Dipteryx alata'' Vogel — Baru
*'' Dipteryx charapilla'' (J.F.Macbr.) Ducke was synonymised to ''D. rosea'' in 1989, but is now recognised as an independent species again.
*'' Dipteryx ferrea'' (Ducke) Ducke was synonymised to ''D. micrantha'' in 1989, but is now recognised as an independent species again.
*'' Dipteryx lacunifera'' Ducke
*'' Dipteryx magnifica'' (Ducke) Ducke
*'' Dipteryx micrantha'' Harms
*''Dipteryx odorata'' (Aubl.) Forsyth f. — Tonka Bean
*'' Dipteryx oleifera'' Benth. — Almendro
*'' Dipteryx polyphylla'' Huber
*'' Dipteryx punctata'' (S.F.Blake) Amshoff
*'' Dipteryx rosea'' Spruce ex Benth.
*'' Dipteryx tetraphylla'' Benth.

References

Further reading

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Dipterygeae
Fabaceae genera
Flora of Southern America