Diplocynodon Darwini
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''Diplocynodon'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
eusuchian Eusuchia is a clade of neosuchian crocodylomorphs that first appeared in the Early Cretaceous, which includes modern crocodilians. Along with Dyrosauridae and Sebecosuchia, they were the only crocodyliformes who survived the K-Pg extinction. ...
, either an
alligatoroid Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members more c ...
crocodilian Crocodilia () is an Order (biology), order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorp ...
or a stem-group crocodilian, that lived during the
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
to
Middle Miocene The Middle Miocene is a sub-epoch of the Miocene epoch (geology), epoch made up of two Stage (stratigraphy), stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0. ...
in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. Some species may have reached lengths of , while others probably did not exceed . They are almost exclusively found in freshwater environments. The various species are thought to have been opportunistic aquatic predators. In the nineteenth century, ''D. steineri'' was named from
Styria Styria ( ; ; ; ) is an Austrian Federal states of Austria, state in the southeast of the country. With an area of approximately , Styria is Austria's second largest state, after Lower Austria. It is bordered to the south by Slovenia, and cloc ...
,
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
and ''D. styriacus'' was named from
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
and
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
. A third Austrian species, ''Enneodon ungeri'', was placed in its own genus. The Austrian and French species of ''Diplocynodon'' were synonymized with ''E. ungeri'' in 2011, and because the name ''Diplocynodon'' has priority over ''Enneodon'', the species is now called ''D. ungeri''. Other
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
have recently been found to be
synonymous A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
with ''Diplocynodon''. ''Hispanochampsa muelleri'' of Spain was determined to be synonymous with ''Diplocynodon'' in 2006, and ''Baryphracta deponaie'' of Germany was confirmed to be synonymous with ''Diplocynodon'' in 2012. Well preserved specimens have been found in the
Messel Pit The Messel Formation is a Formation (geology), geologic formation in Hesse, central Germany, dating back to the Eocene Epoch (geology), epoch (about 47 Ma). Its geographic range is restricted to the Messel pit. There it unconformably overlie ...
and the
Geiseltal The Geisel valley () is a valley in Saxony-Anhalt, Germany, situated west of Merseburg, Saalekreis district. It is named after the River Geisel which rises in Mücheln and is a tributary of the Saale, just under long. Its main settlements are ...
lignite Lignite (derived from Latin ''lignum'' meaning 'wood'), often referred to as brown coal, is a soft, brown, combustible sedimentary rock formed from naturally compressed peat. It has a carbon content around 25–35% and is considered the lowest ...
deposit in
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. Most articulated ''Diplocynodon'' specimens from these localities contain
gastroliths A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In oth ...
. In the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
epoch, the German sites were either a swampy freshwater lake (Messel Pit) or a peat bog swamp (Geiseltal).


Species

*Locality and/or horizon of the type specimen.


Phylogeny

''Diplocynodon'' is one of the basal-most members of the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Alligatoroidea Alligatoroidea is one of three superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous period, and consists of the alligators and caimans, as well as extinct members m ...
. ''Diplocynodon's'' placement within Alligatoroidea can be shown in the
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below, based on a 2018
tip dating Tip dating is a technique used in molecular dating that allows the inference of time-calibrated phylogenetic trees. Its defining feature is that it uses the ages of the samples to provide time information for the analysis, in contrast with traditio ...
study by Lee & Yates that simultaneously used morphological, molecular (
DNA sequencing DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, thymine, cytosine, and guanine. The ...
), and
stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithost ...
(
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
age) data. Below is a more detailed
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
of ''Diplocynodon'': In a 2025 study, Jules D. Walter and colleagues argue that many character states previously thought to be diagnostic for alligatoroids were actually much more widespread. In their analysis several genera traditionally viewed as basal alligatoroids, among them ''Diplocynodon'', were found to not only fall outside of Alligatoroidea but to not even be true crocodilians, instead representing derived non-crocodilian eusuchians.


Palaeobiology

Osteohistological analysis of ''D. hantoniensis'' suggests that it had a similar pattern of growth to the modern American alligator, exhibiting a determinate, seasonally-controlled rate of growth. Additionally, ''D. hantoniensis'' was allometrically very similar to American alligators, with the femoral length and femoral circumference scaling in a similar fashion in both species.
CT scans A computed tomography scan (CT scan), formerly called computed axial tomography scan (CAT scan), is a medical imaging technique used to obtain detailed internal images of the body. The personnel that perform CT scans are called radiographers or ...
of ''D. tormis'' reconstruct internal structures of its snout and a portion of its brain, with implications for its sensory and cognitive capabilities. The relative sizes of the
olfactory bulbs The olfactory bulb (Latin: ''bulbus olfactorius'') is a grey matter, neural structure of the vertebrate forebrain involved in olfaction, the sense of odor, smell. It sends olfactory information to be further processed in the amygdala, the orbitof ...
overlap with alligatoroids and fall short of true crocodiles, adding to anatomical evidence for a closer relationship to alligatoroids. The optic lobes and reptile encephalization quotient are proportionally smaller than other medium-sized crocodilians, though this could be influenced by incomplete preservation at the back of the skull.


Palaeoecology

Based on skull shape analysis of the crocodylians known from the
Lutetian The Lutetian is, in the geologic timescale, a stage (stratigraphy), stage or age (geology), age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it ...
site of Geiseltal, ''Diplocynodon'' was a small generalist carnivore that partitioned its resources with the larger ''
Asiatosuchus ''Asiatosuchus'' is an extinct genus of crocodyloid crocodilians that lived in Eurasia during the Paleogene. Many Paleogene crocodilians from Europe and Asia have been attributed to ''Asiatosuchus'' since the genus was named in 1940. These speci ...
''. According to enamel δ13C values from specimens from the
Late Oligocene The Chattian is, in the geologic timescale The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the pro ...
site of Enspel, the ''Diplocynodon'' living in the palaeoenvironment fed primarily on aquatic vertebrates.


References

* ''Fossils'' (Smithsonian Handbooks) by David Ward (Page 243)


External links


DiscoverGeo Science
{{Taxonbar, from=Q13797209, from2=Q1254623, from3=Q41164376, from4=Q41164378, from5=Q41164384, from6=Q41164387, from7=Q41164391, from8=Q41164396, from9=Q20717785 Alligatoroidea Paleocene crocodylomorphs Eocene crocodylomorphs Oligocene crocodylomorphs Miocene crocodylomorphs Miocene genus extinctions Paleocene reptiles of Europe Paleocene first appearances Eocene reptiles of Europe Oligocene reptiles of Europe Miocene reptiles of Europe Prehistoric pseudosuchian genera Fossils of Austria Fossil taxa described in 1847