Dikdik
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A dik-dik is the name for any of four species of small
antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Madoqua'', which live in the bushlands of eastern and
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
. Dik-diks stand about at the shoulder, are long, weigh and can live for up to 10 years. Dik-diks are named for the alarm calls of the females. In addition to the females' alarm call, both the male and female make a shrill, whistling sound. These calls may alert other animals to predators.


Name

The name ''dik-dik'' comes from an
onomatopoeia Onomatopoeia (or rarely echoism) is a type of word, or the process of creating a word, that phonetics, phonetically imitates, resembles, or suggests the sound that it describes. Common onomatopoeias in English include animal noises such as Oin ...
of the repetitive ''dik'' sound female dik-diks whistle through their long, tubular snouts when they feel threatened.


Physical characteristics

Female dik-diks are somewhat larger than males. The males have
horn Horn may refer to: Common uses * Horn (acoustic), a tapered sound guide ** Horn antenna ** Horn loudspeaker ** Vehicle horn ** Train horn *Horn (anatomy), a pointed, bony projection on the head of various animals * Horn (instrument), a family ...
s, which are small (about ), slanted backwards and longitudinally grooved. The hair on the crown forms an upright tuft that sometimes partially conceals the short, ribbed horns of the male. The upper body is gray-brown, while the lower parts of the body, including the legs, belly, crest, and flanks, are tan. A bare black spot below the inside corner of each eye contains a
preorbital gland The preorbital gland is a paired exocrine gland found in many species of artiodactyls, which is homologous to the lacrimal gland found in humans. These glands are trenchlike slits of dark blue to black, nearly bare skin extending from the media ...
that produces a dark, sticky secretion. Dik-diks insert grass stems and twigs into the gland to scent-mark their territories. Perhaps to prevent overheating, dik-diks (especially Guenther's dik-diks) have elongated snouts with bellows-like muscles through which blood is pumped. Airflow and subsequent evaporation cools this blood before it is recirculated to the body. However, this panting is only implemented in extreme conditions; dik-diks can tolerate air temperatures of up to .


Adaptations for desert environments

Dik-diks have special physiological adaptations to help them survive in arid environments. For instance, dik-diks have a lower density of sweat glands compared to other animals such as cattle. Similarly, in more arid environments, dik-diks can concentrate their urine. These adaptations help dik-diks preserve body water. Because of their small body size, dik-diks are predicted to have among the highest metabolic rates and highest energy requirement per kilogram of all
ruminants Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principa ...
. However, dik-diks have a lower metabolic rate than would be predicted for their size as a physiological adaptation to heat and aridity.


Habitat

Dik-diks live in shrublands and savannas of eastern Africa. Dik-diks seek
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s with a plentiful supply of
edible An edible item is any item that is safe for humans to eat. "Edible" is differentiated from " eatable" because it does not indicate how an item tastes, only whether it is fit to be eaten. Nonpoisonous items found in nature – such as some mushroo ...
plants such as
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s. Dik-diks may live in places as varied as dense forest or open plain, but they require good cover and not too much tall grass. They usually live in pairs in territories of about . The territories are often in low, shrubby bushes (sometimes along dry, rocky streambeds) with plenty of cover. Dik-diks, with their dusty colored coat, are able to blend in with their surroundings. Dik-diks have an established series of runways through and around the borders of their territories that are used when they feel threatened.


Diet

Dik-diks are
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
s. Their diet mainly consists of
foliage A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, f ...
, shoots,
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
and
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...
, but little or no grass. They receive sufficient amounts of water from their food, which makes drinking unnecessary. Like all even-toed ungulates, they digest their food with the aid of micro-organisms in their four-chambered stomachs. After initial digestion, the food is repeatedly eructated and rechewed, a process known also as rumination, or ' chewing the cud'. Dik-diks' tapering heads may help them eat the leaves between the spines on
acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
trees, and feed while still keeping their head high to detect
predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...
s.


Reproduction

Dik-diks are monogamous, and conflicts between territorial neighbors are rare. When they occur, the males from each territory dash at each other, either stop short or make head-to-head contact, then back off for another round, with head crests erected. Males mark their territories with dung piles, and cover the females' dung with their own. One suggestion for monogamy in dik-diks is that it may be an evolutionary response to predation; surrounded by predators, it is dangerous to explore, looking for new partners.''
National Geographic ''National Geographic'' (formerly ''The National Geographic Magazine'', sometimes branded as ''Nat Geo'') is an American monthly magazine published by National Geographic Partners. The magazine was founded in 1888 as a scholarly journal, nine ...
'' "Earth Almanac", June 1996
Pairs spend about 64% of their time together. Males, but not females, will attempt to initiate extra-pair mating if an opportunity arises. Females are sexually mature at six months and males at 12 months. The female gestates for 169 to 174 days and bears a single calf. This happens up to twice a year (at the start and finish of the rainy season). Unlike other ruminants which are born forefeet first, the dik-dik is born nose first, with its forelegs laid back alongside its body. Females weigh about at birth, while males weigh . The mother lactates for six weeks, feeding her calf for no longer than a few minutes at a time. The survival rate for young dik-diks is 50%. The young stay concealed for a time after birth, but grow quickly and reach full size by seven months. At that age, the young are forced to leave their parents' territory. The fathers run the sons off the territory and the mothers run off the daughters.


Predators

Dik-diks are hunted by
leopards The leopard (''Panthera pardus'') is one of the five extant cat species in the genus ''Panthera''. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of with a ...
,
caracal The caracal (''Caracal caracal'') () is a medium-sized Felidae, wild cat native to Africa, the Middle East, Central Asia, and arid areas of Pakistan and northwestern India. It is characterised by a robust build, long legs, a short face, long ...
s,
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...
s,
hyena Hyenas or hyaenas ( ; from Ancient Greek , ) are feliform carnivoran mammals belonging to the family Hyaenidae (). With just four extant species (each in its own genus), it is the fifth-smallest family in the order Carnivora and one of the sma ...
s,
wild dog A free-ranging dog is a dog that is not confined to a yard or house. Free-ranging dogs include street dogs, village dogs, stray dogs, feral dogs, etc., and may be owned or unowned. The global dog population is estimated to be 900 million, of whi ...
s and humans. Other predators include
monitor lizard Monitor lizards are lizards in the genus ''Varanus,'' the only extant genus in the family Varanidae. They are native to Africa, Asia, and Oceania, and West African Nile monitor, one species is also found in south America as an invasive species. A ...
s,
cheetah The cheetah (''Acinonyx jubatus'') is a large Felidae, cat and the Fastest animals, fastest land animal. It has a tawny to creamy white or pale buff fur that is marked with evenly spaced, solid black spots. The head is small and rounded, wit ...
s,
jackal Jackals are Canidae, canids native to Africa and Eurasia. While the word has historically been used for many canines of the subtribe Canina (subtribe), canina, in modern use it most commonly refers to three species: the closely related black-b ...
s,
baboon Baboons are primates comprising the biology, genus ''Papio'', one of the 23 genera of Old World monkeys, in the family Cercopithecidae. There are six species of baboon: the hamadryas baboon, the Guinea baboon, the olive baboon, the yellow ba ...
s,
eagle Eagle is the common name for the golden eagle, bald eagle, and other birds of prey in the family of the Accipitridae. Eagles belong to several groups of Genus, genera, some of which are closely related. True eagles comprise the genus ''Aquila ( ...
s,
hawk Hawks are birds of prey of the family Accipitridae. They are very widely distributed and are found on all continents, except Antarctica. The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks, and others. This ...
s and pythons. Dik-diks' adaptations to predation include excellent eyesight, the ability to reach speeds up to , and high birth rates.


Species

The four species of dik-dik are: * ''Madoqua guntheri''
Thomas Thomas may refer to: People * List of people with given name Thomas * Thomas (name) * Thomas (surname) * Saint Thomas (disambiguation) * Thomas Aquinas (1225–1274) Italian Dominican friar, philosopher, and Doctor of the Church * Thomas the A ...
, 1894
Günther's dik-dik * ''M. kirkii'' ( Günther, 1880)Kirk's dik-dik * ''M. piacentinii'' Drake-Brockman, 1911Silver dik-dik * ''M. saltiana'' ( de Blainville, 1816)
Salt's dik-dik Salt's dik-dik (''Madoqua saltiana'') is a small antelope found in semidesert, bushland, and thickets in the Horn of Africa, but marginally also in northern Kenya and eastern Sudan. It is named after Henry Salt, who was the first European to ack ...


References

{{Taxonbar, from=Q216263   Mammals described in 1837