In terrestrial
vertebrate
Vertebrates () comprise all animal taxon, taxa within the subphylum Vertebrata () (chordates with vertebral column, backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the ...
s, digitigrade () locomotion is walking or running on the toes (from the Latin ''digitus'', 'finger', and ''gradior'', 'walk'). A digitigrade animal is one that stands or walks with its toes (metatarsals) touching the ground, and the rest of its foot lifted. Digitigrades include walking birds (what many assume to be bird knees are actually
ankle
The ankle, or the talocrural region, or the jumping bone (informal) is the area where the foot and the leg meet. The ankle includes three joints: the ankle joint proper or talocrural joint, the subtalar joint, and the inferior tibiofibular ...
s), cats, dogs, and many other
mammals, but not
plantigrades or
unguligrades. Digitigrades generally move more quickly and quietly than other animals.

There are anatomical differences between the
limbs of plantigrades, like humans, and both unguligrade and digitigrade limbs. Digitigrade and unguligrade animals have relatively long
carpals and
tarsals, and the bones which would correspond to the human ankle are thus set much higher in the limb than in a human. In a digitigrade animal, this effectively lengthens the foot, so much so that what are often thought of as a digitigrade animal's "hands" and "feet" correspond only to what would be the
bones of the human finger or toe.
Humans usually walk with the
soles
Sole may refer to:
* Sole (foot), the bottom of the foot
* Sole (shoe), the bottom supporting member of the shoe
Business
* Sole proprietorship, a business which legally has no separate existence from its owner
* Corporation sole, in English la ...
of their feet on the ground, in plantigrade locomotion. In contrast, digitigrade animals walk on their
distal and
intermediate phalanges
The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bon ...
. Digitigrade locomotion is responsible for the distinctive hooked shape of dog legs.
Unguligrade animals, such as horses and cattle, walk only on the distal-most tips of their digits, while in digitigrade animals, more than one segment of the digit makes contact with the ground, either directly (as in birds) or via paw-pads (as in dogs).
Examples
*
Mesonychids
*
Dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
(digitigrade and semi-digitigrade)
**
Birds
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
(except for
loons and
grebes
Grebes () are aquatic diving birds in the order (biology), order Podicipediformes . Grebes are widely distributed freshwater birds, with some species also found in sea, marine habitats during Bird migration, migration and winter. Some flightless ...
which are plantigrade)
*
Pigs (semi-digitigrade)
*
Hippos (semi-digitigrade)
* ''
Pakicetus
''Pakicetus'' is an extinct genus of amphibious cetacean of the family Pakicetidae, which was endemic to Pakistan during the Eocene, about 50 million years ago. It was a wolf-like animal, about to long, and lived in and around water where i ...
''
* ''
Indohyus''
*
Thylacine
The thylacine ( , or , also ) (''Thylacinus cynocephalus'') is an extinct carnivorous marsupial that was native to the Australian mainland and the islands of Tasmania and New Guinea. The last known live animal was captured in 1930 in Tas ...
*
Cats
The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
*
Hyena
Hyenas, or hyaenas (from Ancient Greek , ), are feliform carnivoran mammals of the family Hyaenidae . With only four extant species (each in its own genus), it is the fifth-smallest family in the Carnivora and one of the smallest in the clas ...
s
*
Mongooses
*
Canids
Canidae (; from Latin, ''canis'', "dog") is a family (biology), biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamily, ...
*
Elephants
Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae ...
(semi-digitigrade)
APPEARANCE/ MORPHOLOGY: LEGS, SPINE AND TRACKS with literature reports for the Asian Elephant - Elephas maximus (editorial comment)
/ref>
* Capybaras (semi-digitigrade)
* '' Trucidocynodon'' (semi-digitigrade at least in the forelimbs)
References
External links
Yes, the Shin Bone Is Connected to the Ankle Bone
{{locomotion
Terrestrial locomotion