''Digitalis thapsi'', which has been called mullein foxglove in the US, is a

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

in the genus ''

Digitalis ''Digitalis'' ( or ) is a genus of about 20 species of herbaceous perennial plants, shrubs, and Biennial plant, biennials, commonly called foxgloves. ''Digitalis'' is native to Europe, Western Asia, and northwestern Africa. The flowers are ...

'' that is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...

, where it occurs in eastern

Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...

and central and western

Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...

. It is of commercial importance as an

ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

. Hybrids with ''D. purpurea'' have proved successful and are fertile.

Names

Missouri Botanical Garden The Missouri Botanical Garden is a botanical garden located at 4344 Shaw Boulevard in St. Louis, Missouri. It is also known informally as Shaw's Garden for founder and philanthropy, philanthropist Henry Shaw (philanthropist), Henry Shaw. I ...

website calls it mullein foxglove. Local

vernacular name Vernacular is the ordinary, informal, spoken form of language, particularly when perceived as having lower social status or less prestige than standard language, which is more codified, institutionally promoted, literary, or formal. More n ...

s recorded for this species are ''abeloura'', ''abeloura-amarelada'', ''aboleira'', ''aveloeira'', ''dedaleira'', ''luvas de Santa María'', ''pegajo'' and ''rabo de raposa'' in Portuguese, whereas in

Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many countries in the Americas **Spanish cuisine **Spanish history **Spanish culture ...

the most common name is ''dedalera'', followed by ''viloria'', ''viluria'', ''giloria'', ''dedales'' and ''mataperla'', but it has also been called ''abiloria'', ''abiluria'', ''abortones'', ''bacera'', ''beleño'', ''biloria'', ''biluria'', ''campanilla'', ''cascante'', ''cascaor'', ''chupadera'', ''chupamieles'', ''chupera'', ''cohete'', ''cohetera'', ''cohetes'', ''deales'', ''dedales de niño'', ''dedales purpúreos'', ''dediles'', ''digital'', ''digitalis'', ''emborrachacabras'', ''estallones'', ''goldaperra'', ''guadalperra'', ''gualdaperra'', ''guardaperros'', ''hueltaperra'', ''mata de lagartija'', ''mataperros'', ''rabera'', ''ravera'', ''raéra'', ''restalladera'', ''restralleti'', ''restrallos'', ''sabia'', ''tuercecuellos'' and ''vueltaperra''.

Taxonomy

It was first described in the modern taxonomic system by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in 1763. The genus ''

'' was formerly assigned to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Scrophulariaceae The Scrophulariaceae are a family of flowering plants, commonly known as the figwort family. The plants are annual and perennial herbs, as well as shrubs. Flowers have bilateral (zygomorphic) or rarely radial (actinomorphic) symmetry. The Scr ...

, but it is now considered to belong within the

Plantaginaceae Plantaginaceae, the plantain family or veronica family, is a large, diverse family (biology), family of flowering plants in the order Lamiales that includes common flowers such as Antirrhinum, snapdragon and Digitalis, foxglove. It is unrelated ...

Hybrids

Joseph Gottlieb Kölreuter, a German botanist, observed that ''D. thapsi'' had many characteristics of ''D. purpurea'' after four to five generations of cultivation, and that the former became indistinguishable from the latter, a report that was considered "probably" trustworthy by

Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...

. ''D. thapsi'' and ''D. purpurea''

hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two diff ...

s are generally fertile. In many areas of Spain and Portugal populations of ''D. thapsi'' and ''D. purpurea'' meet, and

introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Introg ...

is common. These have been known as ''D. purpurea'' nothosubsp. ''carpetana'' and ''D. minor'' in error in two older Portuguese works (1906, 1913), but this hybrid is now generally known as ''D.'' × ''coutinhoi''.

Description

Habitus

''Digitalis thapsi'' is a

perennial plant In horticulture, the term perennial (''wikt:per-#Prefix, per-'' + ''wikt:-ennial#Suffix, -ennial'', "through the year") is used to differentiate a plant from shorter-lived annual plant, annuals and biennial plant, biennials. It has thus been d ...

. Its yellow-green leaves are ovate to oblong in shape and decurrent, with winged bases. The flowers, stems and leaves are covered with a dense

indumentum In biology, an indumentum (Latin, literally: "garment") is a covering of trichomes (fine "hairs") on a plant or of bristles (rarely scales) of an insect. Plants The indumentum on plants can have a wide variety of functions, including as ...

of tiny hairs, giving them a soft appearance. These yellow-green hairs (

trichome Trichomes (; ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...

s) are exclusively glandular. The entire plant is very sticky (glutinose). It is

caespitose This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary ...

, meaning it grows with its large leaves tightly arranged into one, or a number, of rosettes growing at or just above ground level. In older plants the base becomes woody and highly branched at ground level, so that a single plant has multiple rosettes. The green, round to slightly angular stems grow to (30-)50 to 80(-100)cm in height before they start to flower. The stems are similarly entirely covered in glandular hairs; these are present in two forms: short and subsessile, or much longer, 0.4 to 0.6mm. The

chromosome count Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, re ...

is 2''n''=56.

Leaves

The lower basal leaf blades are 7.5 to 13 cm (exceptionally 19 cm) in length, 2.5 to 5 cm in width, more or less flat but sometimes undulated along the margins, and oblong to elliptic in shape, with a sharp to somewhat sharply tipped apex. The bases of the leaves taper gradually into the petioles. The texture is soft, not leathery. The underside is

rugose Rugose means "wrinkled". It may refer to: * Rugosa, an extinct order of coral, whose rugose shape earned it the name * Rugose, adjectival form of rugae Species with "rugose" in their names * ''Idiosoma nigrum'', more commonly, a black rugose tra ...

-reticular in texture, coloured greenish-yellow and very densely packed with glandular hairs. The leaves in the middle of the rosette are clearly

decurrent ''Decurrent'' (sometimes decurring) is a term used in botany and mycology to describe plant or fungal parts that extend downward. In botany, the term is most often applied to leaf blades that partly wrap or have wings around the stem or petio ...

. The margins are denticulate or somewhat so, rarely subentire.

Flowers

The pink, rarely white, flowers are arranged in an

inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...

in the form of a

raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...

, 15 to 35 cm in length with 15 to 40 individual flowers. The inflorescence has glandular hairs along its shaft, is not stiff, and is secundiflorous, meaning the flowers follow each other in succession. The flowers have a 1 to 2 cm long

pedicel Pedicle or pedicel may refer to: Human anatomy *Pedicle of vertebral arch, the segment between the transverse process and the vertebral body, and is often used as a radiographic marker and entry point in vertebroplasty and kyphoplasty procedures ...

which is curved at the base, and shorter to equal in length to the subtending

bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...

s. The flowers hang somewhat, and are separated by internodes of 8-15mm. The bracts are 12 to 20mm in length and 2 to 3mm broad, glandular and

lanceolate The following terms are used to describe leaf plant morphology, morphology in the description and taxonomy (biology), taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade ...

in shape.

Chemistry

According to Dutch botanist

Herman Boerhaave Herman Boerhaave (, 31 December 1668 – 23 September 1738Underwood, E. Ashworth. "Boerhaave After Three Hundred Years." ''The British Medical Journal'' 4, no. 5634 (1968): 820–25. .) was a Dutch chemist, botanist, Christian humanist, and ph ...

, writing at the turn of the 17th century, ''Digitalis'' species are highly poisonous if directly ingested. All parts are poisonous. In studies, the restriction of

calcium Calcium is a chemical element; it has symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to it ...

ions resulted in

cardenolide A cardenolide is a type of steroid. Many plants contain derivatives, collectively known as cardenolides, including many in the form of cardenolide glycosides (cardenolides that contain structural groups derived from sugars). Cardenolide glycoside ...

accumulation in ''D. thapsi''. Calcium quantity affects the

redox Redox ( , , reduction–oxidation or oxidation–reduction) is a type of chemical reaction in which the oxidation states of the reactants change. Oxidation is the loss of electrons or an increase in the oxidation state, while reduction is t ...

chemical reactions in the cells. Without calcium, changes in

antioxidant Antioxidants are Chemical compound, compounds that inhibit Redox, oxidation, a chemical reaction that can produce Radical (chemistry), free radicals. Autoxidation leads to degradation of organic compounds, including living matter. Antioxidants ...

function were observed and

catalase Catalase is a common enzyme found in nearly all living organisms exposed to oxygen (such as bacteria, plants, and animals) which catalyzes the decomposition of hydrogen peroxide to water and oxygen. It is a very important enzyme in protecting ...

activity was slow. In another study, lack of calcium retarded growth and promoted

digoxin Digoxin (better known as digitalis), sold under the brand name Lanoxin among others, is a medication used to treat various heart disease, heart conditions. Most frequently it is used for atrial fibrillation, atrial flutter, and heart failure. ...

formation.

Manganese sulfate Manganese(II) sulfate usually refers to the inorganic compound with the formula MnSO4·H2O. This pale pink deliquescent solid is a commercially significant manganese(II) salt. Approximately 260,000 tonnes of manganese(II) sulfate were produced wor ...

and

lithium chloride Lithium chloride is a chemical compound with the formula Li Cl. The salt is a typical ionic compound (with certain covalent characteristics), although the small size of the Li+ ion gives rise to properties not seen for other alkali metal chlorid ...

also increased the digoxin concentration, but did not affect growth.

Calcium oxalate Calcium oxalate (in archaic terminology, oxalate of lime) is a calcium salt of oxalic acid with the chemical formula or . It forms hydrates , where ''n'' varies from 1 to 3. Anhydrous and all hydrated forms are colorless or white. The monohydr ...

crystals have also been isolated from ''D. thapsi''.

Similar species

In 1841

Pierre Edmond Boissier Pierre Edmond Boissier (25 May 1810 Geneva – 25 September 1885 Valeyres-sous-Rances) was a Swiss prominent botanist, explorer and mathematician. He was the son of Jacques Boissier (1784-1857) and Caroline Butini (1786-1836), daughter of Pierr ...

considered it the most similar to ''D. mariana'', which he was then describing as a new species, having collected from the same region. In the

dichotomous key In phylogenetics, a single-access key (also called dichotomous key, sequential key, analytical key, or pathway key) is an identification key where the sequence and structure of identification steps is fixed by the author of the key. At each point i ...

in the ''

Flora Iberica ''Flora Iberica: Plantas vasculares de la Península Ibérica e Islas Baleares'' ("Vascular plants of the Iberian Peninsula and Balearic Islands") is a Spanish book series containing identification keys, descriptions, and illustrations of pterido ...

'', it is keyed out to ''D. mariana'', ''D. minor'' and ''D. purpurea''. ''D. thapsi'' is distinguished from these three other similar species by being the only very sticky species, being completely covered in relatively long, yellowish, glandular hairs. It is also the only species in which the leaves in the middle of the rosette are clearly decurrent. ''D. minor'', an endemic of the

Balearic Islands The Balearic Islands are an archipelago in the western Mediterranean Sea, near the eastern coast of the Iberian Peninsula. The archipelago forms a Provinces of Spain, province and Autonomous communities of Spain, autonomous community of Spain, ...

, is not

sympatric In biology, two closely related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter each other. An initially interbreeding population that splits into two or more distinct spe ...

with ''D. thapsi''.

Distribution

''Digitalis thapsi'' is an

species of the

, occurring in both

and

. It grows in the mountains and rocky plains of the central and central-western parts of the Iberian Peninsula, especially in the

Sistema Central The Central System, Spanish language, Spanish and , is one of the main systems of mountain ranges in the Iberian Peninsula. The 2,592 m high Pico Almanzor is its highest summit. The Central System is located just north of the 40th parallel north, ...

and its extensions between the rivers of the

Douro The Douro (, , , ; ; ) is the largest river of the Iberian Peninsula by discharge. It rises near Duruelo de la Sierra in the Spanish Soria Province, province of Soria, meanders briefly south, then flows generally west through the northern par ...

and

Tagus The Tagus ( ; ; ) is the longest river in the Iberian Peninsula. The river rises in the Montes Universales between Cuenca and Teruel, in mid-eastern Spain, flows , generally westward, and empties into the Atlantic Ocean in Lisbon. Name T ...

. It is widespread in Spain, occurring in the western and central parts of the country. It does not occur on the

. It occurs in the provinces of

Ávila Ávila ( , , ) is a Spanish city located in the autonomous community of Castile and León. It is the capital and most populated municipality of the Province of Ávila. It lies on the right bank of the Adaja river. Located more than 1,130 m a ...

Badajoz Badajoz is the capital of the Province of Badajoz in the autonomous communities of Spain, autonomous community of Extremadura, Spain. It is situated close to the Portugal, Portuguese Portugal–Spain border, border, on the left bank of the river ...

, Cáceres,

Córdoba Córdoba most commonly refers to: * Córdoba, Spain, a major city in southern Spain and formerly the imperial capital of Islamic Spain * Córdoba, Argentina, the second largest city in Argentina and the capital of Córdoba Province Córdoba or Cord ...

Guadalajara Guadalajara ( ; ) is the capital and the most populous city in the western Mexican List of states of Mexico, state of Jalisco, as well as the most densely populated municipality in Jalisco. According to the 2020 census, the city has a population ...

Madrid Madrid ( ; ) is the capital and List of largest cities in Spain, most populous municipality of Spain. It has almost 3.5 million inhabitants and a Madrid metropolitan area, metropolitan area population of approximately 7 million. It i ...

Salamanca Salamanca () is a Municipality of Spain, municipality and city in Spain, capital of the Province of Salamanca, province of the same name, located in the autonomous community of Castile and León. It is located in the Campo Charro comarca, in the ...

Segovia Segovia ( , , ) is a city in the autonomous communities of Spain, autonomous community of Castile and León, Spain. It is the capital and most populated municipality of the Province of Segovia. Segovia is located in the Meseta central, Inner Pl ...

Soria Soria () is a municipality and a Spanish city, located on the Douro river in the east of the autonomous community of Castile and León and capital of the province of Soria. Its population is 38,881 ( INE, 2017), 43.7% of the provincial populatio ...

Toledo Toledo most commonly refers to: * Toledo, Spain, a city in Spain * Province of Toledo, Spain * Toledo, Ohio, a city in the United States Toledo may also refer to: Places Belize * Toledo District * Toledo Settlement Bolivia * Toledo, Or ...

and Zamora. It has incidentally been found in the

province of Burgos The province of Burgos is a Provinces of Spain, province of northern Spain, in the northeastern part of the Autonomous communities of Spain, autonomous community of Castile-Leon, Castile and León. It is bordered by the provinces of Palencia (p ...

. In

Andalucía Andalusia ( , ; , ) is the southernmost autonomous communities of Spain, autonomous community in Peninsular Spain, located in the south of the Iberian Peninsula, in southwestern Europe. It is the most populous and the second-largest autonomou ...

it is only found in the north of the region, in the western

Sierra Morena The Sierra Morena is one of the main systems of mountain ranges in Spain. It stretches for 450 kilometres from east to west across the south of the Iberian Peninsula, forming the southern border of the ''Meseta Central'' plateau and providi ...

, where it is uncommon; it has been collected more often in the mountains of central Spain. In Portugal it primarily occurs in the northern and central interior, more precisely in the districts of

Bragança Bragança may refer to: People *Jaime Celestino Dias Bragança, a Portuguese footballer Politics and History *House of Bragança - A Portuguese Royal House *Duke of Bragança - A Portuguese noble, and later royal, title * Bragança - One of the m ...

Vila Real Vila Real () is the capital and largest city of the Vila Real District, in the Norte, Portugal, North region, Portugal. It is also the seat of the Douro (intermunicipal community), Douro Intermunicipal communities of Portugal, intermunicipal comm ...

Guarda Guarda may refer to: * Guarda, Portugal, city in Portugal * Guarda District, the district containing that city * Roman Catholic Diocese of Guarda, Portugal, the diocese containing that city * Guarda, Switzerland, municipality in Grisons, Switzerlan ...

Viseu Viseu () is a city and municipality in the Centro Region of Portugal and the capital of the Viseu District, district of the same name, with a population of 100,105 inhabitants in the entire municipality, and center of the Viseu Dão Lafões Interm ...

, Castelo Branco, Portalegre and

Évora Évora ( , ), officially the Very Noble and Ever Loyal City of Évora (), is a city and a municipalities of Portugal, municipality in Portugal. It has 53,591 inhabitants (2021), in an area of . It is the historic capital of the Alentejo reg ...

. With less frequency, it has also been found in the

Porto Porto (), also known in English language, English as Oporto, is the List of cities in Portugal, second largest city in Portugal, after Lisbon. It is the capital of the Porto District and one of the Iberian Peninsula's major urban areas. Porto c ...

, Aveiro,

Coimbra Coimbra (, also , , or ), officially the City of Coimbra (), is a city and a concelho, municipality in Portugal. The population of the municipality at the 2021 census was 140,796, in an area of . The fourth-largest agglomerated urban area in Po ...

, Santarém, Beja and Faro Districts.

Ecology

''Digitalis thapsi'' grows in acidic soils. It grows on rocky ground and on slopes. It prefers sunny, open and disturbed places. It generally grows on soils derived from

granite Granite ( ) is a coarse-grained (phanerite, phaneritic) intrusive rock, intrusive igneous rock composed mostly of quartz, alkali feldspar, and plagioclase. It forms from magma with a high content of silica and alkali metal oxides that slowly coo ...

, but also sometimes on

quartzite Quartzite is a hard, non- foliated metamorphic rock that was originally pure quartz sandstone.Essentials of Geology, 3rd Edition, Stephen Marshak, p 182 Sandstone is converted into quartzite through heating and pressure usually related to tecton ...

and

schist Schist ( ) is a medium-grained metamorphic rock generally derived from fine-grained sedimentary rock, like shale. It shows pronounced ''schistosity'' (named for the rock). This means that the rock is composed of mineral grains easily seen with a l ...

. It grows at (rarely 100-) 500–1500 metres in altitude. The specific phytosociological suballiance and alliance wherein it occurs is called 'Rumici indurati-Dianthion lusitani', which is in the order 'Phagnalo saxatilis-Rumicetalia indurati', in the class 'Phagnalo saxatilis-Rumicetea indurati'. It is a characteristic species of this phytocoenosis. In this

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

it occurs together with these following characteristic species: ''Antirrhinum graniticum'' subsp. ''graniticum'', '' Antirrhinum rupestris'', '' Arrhenatherum fernandesii'', '' Biscutella bilbilitana'', ''Centaurea monticola'' subsp. ''citricolor'', '' Centaurea pinnata'', '' Coincya leptocarpa'', '' Coincya longirostra'', ''Coincya pseudoerucastrum'' subsp. ''pseudoerucastrum'', '' Coincya rupestris'', ''Conopodium bunioides'' subsp. ''aranii'', ''Conopodium majus'' subsp. ''marizianum'', '' Crepis oporinoides'', ''Dianthus crassipes'' subsp. ''crassipes'', ''Dianthus crassipes'' subsp. ''sereneanus'', '' Dianthus lusitanus'', '' Digitalis mariana'' (both subspecies), ''Digitalis purpurea'' subsp. ''toletana'', '' Erodium mouretti'', ''Erysimum linifolium'' subsp. ''lagascae'', '' Festuca duriotagana'', ''Jasione sessiliflora'' subsp. ''tomentosa'', '' Scrophularia oxyrhincha'', '' Scrophularia sublyrata'', ''Sedum hirsutum'' subsp. ''baeticum'', '' Silene marizii'', ''

Silene × montistellensis ''Silene'' is a genus of flowering plants in the family Caryophyllaceae. Containing nearly 900 species, it is the largest genus in the family. Common names include campion and catchfly. Many ''Silene'' species are widely distributed, particularl ...

'' and ''Verbascum rotundifolium'' subsp. ''castellanum''. In its native lands it flowers from June to July. In cultivation it flowers in late spring to early summer. A study of the fruiting activity in ''D. thapsi'' showed that production, size and number of seeds were interrelated. Analysis has shown that seed weight is not related to the length of the

cotyledon A cotyledon ( ; ; "a cavity, small cup, any cup-shaped hollow", gen. (), ) is a "seed leaf" – a significant part of the embryo within the seed of a plant – and is formally defined as "the embryonic leaf in seed-bearing plants, one or mor ...

, and that the length of the fifth leaf can serve as a useful factor in determining the

genetic variability Genetic variability is either the presence of, or the generation of, genetic differences. It is defined as "the formation of individuals differing in genotype, or the presence of genotypically different individuals, in contrast to environmentally ...

among specimens.

Uses

Agriculture

It is known that

livestock Livestock are the Domestication, domesticated animals that are raised in an Agriculture, agricultural setting to provide labour and produce diversified products for consumption such as meat, Egg as food, eggs, milk, fur, leather, and wool. The t ...

avoid eating this plant, but an exception are goats, which will eat the leaves and flowers in times of shortage. Nonetheless, the plant is poisonous for them, causing stupor and paralysis -this was apparently once not an uncommon occurrence in Spain, and has spawned a number of idioms.

Cultivation

The species is used as a perennial,

. It is low-growing, so is used in the border. It grows well in half-shaded to sun-exposed areas in temperate regions. It prefers organically rich, reasonably well-drained, acidic soils. In Missouri it is said to require constantly moist soil. In British gardens moderate watering is required; the plant has an average drought tolerance, and is adaptable to dry shady areas. It is tolerant of deer. Many gardeners remove the spent flower spikes soon after bloom, not allowing it to go to seed. Besides normal cultivation, ''D. thapsi'' has been propagated using

explant culture In biology, explant culture is a technique to organotypically culture cells from a piece or pieces of tissue or organ removed from a plant or animal. The term ''explant'' can be applied to samples obtained from any part of the organism. The extract ...

, a technique employed by isolating and harvesting

meristem In cell biology, the meristem is a structure composed of specialized tissue found in plants, consisting of stem cells, known as meristematic cells, which are undifferentiated cells capable of continuous cellular division. These meristematic c ...

cells from pieces of tissue.

Auxin Auxins (plural of auxin ) are a class of plant hormones (or plant-growth regulators) with some morphogen-like characteristics. Auxins play a cardinal role in coordination of many growth and behavioral processes in plant life cycles and are essent ...

s NAA, 2,4-D and IAA alone or combined with BA produced a

callus A callus (: calluses) is an area of thickened and sometimes hardened skin that forms as a response to repeated friction, pressure, or other irritation. Since repeated contact is required, calluses are most often found on the feet and hands, b ...

. NAA caused root formation and BA shoot formation. NAA and BA combined induced organ generation more effectively. Plantlets obtained this way had a survival rate of 70%. A

cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...

called 'Spanish Peaks' with raspberry-rose-coloured flowers and a compact habitus has been available in the US.

Traditional

In Spain the traditional uses have practically been abandoned. Recorded traditional uses are often identical to that those of ''D. purpurea''; when questioned many informants in Salamanca believed that these were in fact the same species. It is generally thought that the difference in flower colour is due to some characteristic of the soil. Local people are aware it is poisonous. It was once locally used in

herbalism Herbal medicine (also called herbalism, phytomedicine or phytotherapy) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. Scientific evidence for the effectiveness of many herbal treatments ...

as a cardiac tonic in many places. Other folk medicinal uses are local to specific cities or villages: in the hills near Salamanca the leaves were steeped in water to use for a sore throat or a

decoction Decoction is a method of extraction by boiling herbal or plant material (which may include stems, roots, bark and rhizomes) to dissolve the chemicals of the material. It is the most common preparation method in various herbal medicine systems. D ...

used for infected wounds, in a town in Galicia the flowers in water were used to combat inflammations, in the provinces of Zamora and Salamanca boils are said to heal after applying a

poultice A poultice or cataplasm, also called a fomentation, is a soft moist mass, often heated and medicated, that is applied to the skin to reduce inflammation, soothe pain, promote healing, or otherwise treat wounds or ailments. Soft materials like cer ...

of the burnt basal leaves.

Conservation

It is not considered rare in a national or international perspective, but it has been classified as locally '

endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...

' in the

Regional Red List A Regional Red List is a report of the threatened status of species within a certain country or region. It is based on the IUCN Red List of Threatened Species, an inventory of the conservation status of species on a global scale. Regional Red ...

of Andalucía of 2005. The reason for this was that the different collection localities and populations in this region were fragmented and very small.

References

{{Taxonbar, from=Q5276216 thapsi Plants described in 1753 Flora of Portugal Flora of Spain Taxa named by Carl Linnaeus Endemic flora of the Iberian Peninsula