Marine hatchetfishes or deep-sea hatchetfishes are small

deep-sea The deep sea is broadly defined as the ocean depth where light begins to fade, at an approximate depth of or the point of transition from continental shelves to continental slopes. Conditions within the deep sea are a combination of low tempe ...

mesopelagic The mesopelagic zone (Greek language, Greek μέσον, middle), also known as the middle pelagic or twilight zone, is the part of the pelagic zone that lies between the photic epipelagic and the aphotic bathypelagic zones. It is defined by light ...

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

of the

stomiiform Stomiiformes is an order of deep-sea ray-finned fishes of very diverse morphology. It includes, for example, dragonfishes, lightfishes (Gonostomatidae and Phosichthyidae), loosejaws, marine hatchetfishes and viperfishes. The order contains 4 fa ...

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Sternoptychinae. They should not be confused with the

freshwater hatchetfish The freshwater hatchetfish are a family, Gasteropelecidae, of ray-finned fish from South and Central America. The common hatchetfish is the most popular member among fish keeping hobbyists. The family includes three genera: '' Carnegiella'' (f ...

es, which are not particularly closely related

Teleostei Teleostei (; Ancient Greek, Greek ''teleios'' "complete" + ''osteon'' "bone"), members of which are known as teleosts (), is, by far, the largest group of ray-finned fishes (class Actinopterygii), with 96% of all neontology, extant species of f ...

in the

characiform Characiformes is an order of Actinopterygii, ray-finned fish, comprising the characins and their allies. Grouped in 18 recognized families, more than 2000 different species are described, including the well-known piranha and tetras.; Buckup P.A ...

family Gasteropelecidae. (2006): ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' (4th ed.). John Wiley & Sons, Inc. p.209 The

scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

means "''Sternoptyx''-subfamily", from ''

Sternoptyx ''Sternoptyx'' is an oceanic ray-finned fish genus which belongs in the family (biology), family Sternoptychidae. This is the type genus of the Sternoptychidae, as well as the marine hatchetfish subfamily Sternoptychinae. ''Sternoptyx'' have si ...

'' (the

type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...

) + the standard

animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...

family

suffix In linguistics, a suffix is an affix which is placed after the stem of a word. Common examples are case endings, which indicate the grammatical case of nouns and adjectives, and verb endings, which form the conjugation of verbs. Suffixes can ca ...

"-inae". It ultimately derives from

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

''stérnon'' (στέρνον, "breast") + ''ptýx'' (πτύξ, "a fold/crease") +

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

''forma'' ("external form"), the Greek part in reference to the

thorax The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main di ...

shape of marine hatchetfishes.

Description and ecology

Found in

tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...

subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...

and

temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...

waters of the

Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is bounded by the cont ...

and

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or approximately 20% of the water area of Earth#Surface, Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia (continent), ...

s, marine hatchetfishes range in size from ''

Polyipnus danae ''Polyipnus danae'' is a species of ray-finned fish in the genus ''Polyipnus''. It is found in the South China Sea The South China Sea is a marginal sea of the Western Pacific Ocean. It is bounded in the north by South China, in the west by t ...

'' at to the c.-long

giant hatchetfish ''Argyropelecus gigas'', the giant hatchetfish or greater silver hatchetfish, is a marine fish of the genus ''Argyropelecus''. It is found in every ocean except the north Pacific in the mesopelagic zone of tropical and subtropical waters. "Giant" ...

(''Argyropelecus gigas''). They are small

fishes which have

evolved Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...

a peculiar body shape and like their relatives have

bioluminescent Bioluminescence is the emission of light during a chemiluminescence reaction by living organisms. Bioluminescence occurs in multifarious organisms ranging from marine vertebrates and invertebrates, as well as in some fungi, microorganisms inc ...

photophore A photophore is a specialized anatomical structure found in a variety of organisms that emits light through the process of boluminescence. This light may be produced endogenously by the organism itself (symbiotic) or generated through a mut ...

s. The latter allow them to use

counter-illumination Counter-illumination is a method of active camouflage seen in marine animals such as firefly squid and midshipman fish, and in military prototypes, producing light to match their backgrounds in both brightness and wavelength. Marine animals of ...

to escape

predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...

s that lurk in the depths: by matching the light intensity with the light penetrating the water from above, the fish does not appear darker if seen from below. They typically occur at a few hundred meters below the surface, but their entire depth range spans from 50 to 1,500 meters deep. (1998): Sternoptychidae. ''In:'' : ''Encyclopedia of Fishes'': 121. Academic Press, San Diego. The body is deep and

lateral Lateral is a geometric term of location which may also refer to: Biology and healthcare * Lateral (anatomy), a term of location meaning "towards the side" * Lateral cricoarytenoid muscle, an intrinsic muscle of the larynx * Lateral release ( ...

ly extremely compressed, somewhat resembling a

hatchet A hatchet (from the Old French language, Old French , a diminutive form of ''hache'', 'axe' of Germanic origin) is a Tool, single-handed striking tool with a sharp blade on one side used to cut and split wood, and a hammerhead on the other side ...

(with the

being the "blade" and the

caudal peduncle Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

being the "handle"). The

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

Polyipnus ''Polyipnus'' is a genus of oceanic ray-finned fish in the family Sternoptychidae. This is the largest genus of the marine hatchetfishes subfamily Sternoptychinae and indeed of the entire Sternoptychidae. It is not quite as apomorphic as their r ...

'' is rounded, the other two – in particular ''

'' – decidedly angular if seen from the side. Their

pelvis The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...

is rotated to a vertical position. The mouth is located at the tip of the snout and directed almost straight downwards. Their scales are silvery, delicate and easily abraded. In some species, such as the highlight hatchetfish (''

Sternoptyx pseudobscura ''Sternoptyx'' is an oceanic ray-finned fish genus which belongs in the family Sternoptychidae. This is the type genus of the Sternoptychidae, as well as the marine hatchetfish subfamily Sternoptychinae. ''Sternoptyx'' have silvery, high, and ...

''), large sections of the body at the base of the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

and/or

caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

are transparent. They have

perpendicular In geometry, two geometric objects are perpendicular if they intersect at right angles, i.e. at an angle of 90 degrees or π/2 radians. The condition of perpendicularity may be represented graphically using the '' perpendicular symbol'', � ...

spines and blade-like

pterygiophore A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine biology, marine environments, so the fins are not all Homology (biol ...

s in front of the

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

. The

has 11–19

rays Ray or RAY may refer to: Fish * Ray (fish), any cartilaginous fish of the superorder Batoidea * Ray (fish fin anatomy), the bony or horny spine on ray-finned fish Science and mathematics * Half-line (geometry) or ray, half of a line split at an ...

and in some

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

is divided in two parts; almost all have an

adipose fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only b ...

. Their large, sometimes tube-shaped eyes can collect the faintest of light and

focus Focus (: foci or focuses) may refer to: Arts * Focus or Focus Festival, former name of the Adelaide Fringe arts festival in East Australia Film *Focus (2001 film), ''Focus'' (2001 film), a 2001 film based on the Arthur Miller novel *Focus (2015 ...

well on objects both close and far. They are directed somewhat upwards, most conspicuously in the genus ''

Argyropelecus ''Argyropelecus'' is an oceanic Actinopterygii, ray-finned fish genus in the deep sea hatchetfish family (biology), family Sternoptychidae. A collective name is "Marine hatchetfish, silver hatchetfishes", but this can also refer to a species of ...

''. This allows them to discern the

silhouette A silhouette (, ) is the image of a person, animal, object or scene represented as a solid shape of a single colour, usually black, with its edges matching the outline of the subject. The interior of a silhouette is featureless, and the silhouett ...

s of

prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...

moving overhead against the slightly brighter upper waters.

Genera

There are three genera in this

, with some 40

altogether: * ''

'' – silver hatchetfishes (7 species) * ''

'' (32 species) * ''

'' (4 species) Four fossil genera are also known: * '' Eosternoptyx'' (middle-late

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...

) * '' Polyipnoides'' (middle Eocene of

Georgia Georgia most commonly refers to: * Georgia (country), a country in the South Caucasus * Georgia (U.S. state), a state in the southeastern United States Georgia may also refer to: People and fictional characters * Georgia (name), a list of pe ...

) * '' Horbatshia'' (

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

of the

Carpathians The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe and Southeast Europe. Roughly long, it is the third-longest European mountain range after the Urals at and the Scandinavian Mountains ...

) * '' Discosternon'' (

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...

)

Footnotes

{{Taxonbar, from=Q6764289 Sternoptychidae Bioluminescent fish Deep sea fish