''Dasygrammitis'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

fern The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...

s in the family

Polypodiaceae Polypodiaceae is a Family (biology), family of ferns. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), the family includes around 65 genus, genera and an estimated 1,650 species and is placed in the order Polypodiales, suborder ...

, subfamily

Grammitidoideae Grammitidoideae is a subfamily of the fern family Polypodiaceae, whose members are informally known as grammitids. It comprises a clade of about 750 species. They are distributed over higher elevations in both the Old World, Old and New World. T ...

, according to the

Pteridophyte Phylogeny Group The Pteridophyte Phylogeny Group (PPG) is an informal international group of systematic botanists who collaborate to establish on the classification of pteridophytes ( lycophytes and ferns) that reflects knowledge about plant relationships discove ...

classification of 2016 (PPG I). It is known from

Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...

through southeast Asia and the Philippines to Polynesia.

Description

The rhizomes are radially symmetric (without distinct upper and lower surfaces) and bear whorls of stipes, which lack a joint at the point of attachment. The rhizome scales are red-brown, of uniform color, and usually glossy. They have unbranched, red-brown hairs on their edges. Hairs, where present, are unbranched and branched, and brown in color. The leaf blades are usually

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...

in cutting, rarely

pinnatifid Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...

or bipinnatifid, bearing free veins which lack

hydathode A hydathode is a type of pore, commonly found in vascular plants, that secretes water through pores in the epidermis or leaf margin, typically at the tip of a marginal tooth or serration. Hydathodes occur in the leaves of submerged aquatic plant ...

s at their terminus. Sori are borne in two rows beneath pinnae or lobes. The sori are circular to elliptic in shape; the

sporangia A sporangium (from Late Latin, ; : sporangia) is an enclosure in which spores are formed. It can be composed of a unicellular organism, single cell or can be multicellular organism, multicellular. Virtually all plants, fungus, fungi, and many ot ...

either lack hairs or have one to two red-brown hairs.

Taxonomy

The genus was first described by Barbara Parris in 2007 to receive some of the species of the genus ''

Grammitis ''Grammitis'' (dwarf polypody) is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It had formerly been placed in the family Grammitidacea ...

'', which as then circumscribed was artificial and contained a number of disparate lineages. The name "''Dasygrammitis''" means "shaggy ''Grammitis''", referring to the shaggy hairs often present on the frond in this group of former ''Grammitis'' species. Parris initially placed six species in the genus. A phylogenetic study of the grammitids found that ''Dasygrammitis'' was sister to a clade consisting of ''

Calymmodon ''Calymmodon'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group The Pteridophyte Phylogeny Group (PPG) is an informal international group of systematic botanists who colla ...

'', ''

Micropolypodium ''Micropolypodium'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is native to China, Japan, and the Philippines. Species Neotro ...

'', '' Scleroglossum'', '' Tomophyllum'', and '' Xiphopterella''. These six genera share a radially symmetric rhizome, which may be a

synapomorphy In phylogenetics, an apomorphy (or derived trait) is a novel Phenotypic trait, character or character state that has evolution, evolved from its ancestral form (or Plesiomorphy and symplesiomorphy, plesiomorphy). A synapomorphy is an apomorphy sh ...

for this clade (although the trait appears in other grammitid taxa, probably independently).

Species

, the ''Checklist of Ferns and Lycophytes of the World'' accepted the following eight species: *'' Dasygrammitis brevivenosa'' (Alderw.) Parris *'' Dasygrammitis crassifrons'' (Baker) Parris *'' Dasygrammitis fuscata'' (Blume) Parris *''Dasygrammitis kinabaluensis'' Parris, sp. ined. *'' Dasygrammitis malaccana'' (Baker) Parris *''

Dasygrammitis mollicoma ''Dasygrammitis'' is a genus of ferns in the family Polypodiaceae, subfamily Grammitidoideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). It is known from Sri Lanka through southeast Asia and the Philippines ...

'' (Nees & Blume) Parris *'' Dasygrammitis purpurascens'' (Nadeaud) Parris *''Dasygrammitis seramensis'' Parris, sp. ined.

References

{{Taxonbar, from=Q17197627 Polypodiaceae Fern genera