The dara (''Parakuhlia macrophthalmus'') is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

, a

grunt Grunt, grunts or grunting may refer to: Sound and music * Grunting (tennis), in tennis refers to the loud noise, sometimes described as "shrieking" or "screaming", made by some players during their strokes * Death grunt, the death metal singi ...

belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Haemulidae Haemulidae is a family (biology), family of fishes in the order (biology), order Perciformes known commonly as grunts. It is made up of the two subfamily, subfamilies Haemulinae (grunts) and Plectorhynchinae (sweetlips), which contain about 133 ...

. It is native to the

Atlantic The Atlantic Ocean is the second largest of the world's five oceanic divisions, with an area of about . It covers approximately 17% of Earth's surface and about 24% of its water surface area. During the Age of Discovery, it was known for se ...

coast of

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

. It is the only species in the

monospecific In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

''Parakuhlia''.

Description

The dara has an oval, moderately deep and laterally compressed body with a steep head, the profile of the head being , a little concave above the large eyes and ending in a short, blunt snout. The mouth is large, set diagonally and has protrusible jaws. The jaws are equipped with many bands made up of villiform teeth which are also on the vomer but there are no teeth on the palate There is a single

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

but it has a deep notch between the spiny portion and the soft-rayed portion. The dorsal fin contains 11 spines in before the notch and 1 spine and 15 or 16 soft rays behind the notch, the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

has 3 spines and 16 soft rays. The

caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

is slightly emarginated. It is mostly silvery in colour, darker on the back, while the fins are dark yellow. This species attains a maximum total length of , although is more typical.

Distribution

The dara is found in the eastern Atlantic Ocean along the western coast of Africa from Senegal in the north to Angola in the south, it is commonest in the

Gulf of Guinea The Gulf of Guinea (French language, French: ''Golfe de Guinée''; Spanish language, Spanish: ''Golfo de Guinea''; Portuguese language, Portuguese: ''Golfo da Guiné'') is the northeasternmost part of the tropical Atlantic Ocean from Cape Lopez i ...

Habitat and biology

The dara is found at depths between where it is found in inshore waters at rocky coasts and beaches. It is an

oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...

species which forms distinct pairs for

spawning Spawn is the Egg cell, eggs and Spermatozoa, sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of freely releasing eggs and sperm into a body of water (fresh or marine); the physical act is ...

. Almost nothing is known about the biology of this species.

Systematics

The dara was first formally described in 1893 as ''Haemulon macropthalmus'' by the Portuguese naturalist Balthasar Osório with the type locality given as Rolas Island off

São Tomé Island São Tomé Island, at , is the largest island of São Tomé and Príncipe and is home in May 2018 to about 193,380 or 96% of the nation's population. The island is divided into six districts of São Tomé and Príncipe, districts. It is located ...

. In 1913 the

French French may refer to: * Something of, from, or related to France ** French language, which originated in France ** French people, a nation and ethnic group ** French cuisine, cooking traditions and practices Arts and media * The French (band), ...

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Jacques Pellegrin (1873-1944) described a new species and placed it in the new genus ''Parakuhlia'', giving it the

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''boulengeri''. Pellegrin's taxon was later shown to be a

junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

of Osório's ''Haemulon macropthalmus''. ''Parakuhlia'' is monotypic, the dara being the only species in this genus. This species was originally placed in the Flagtail family,

Kuhliidae The flagtails (' or ' in the Hawaiian language) are a family (biology), family (Kuhliidae) of centrarchiform ray-finned fish of the Indo-Pacific area. The family consists of about 12 species in one genus, ''Kuhlia''. Most are euryhaline and often ...

and the generic name ''Parakuhlia'' is a reference to Pellegrin's note that this species is similar in appearance to '' Kuhlia caudovittata'' although it differed in a number of characteristics. The specific name ''marcropthalma'' means large-eyed.

Utilisation

The dara is not considered to be important to commercial fisheries but it is caught by artisanal fisheries using seines, set nets and hook and line. The fish are sold fresh and normally eaten fried or

smoked Smoking is the process of flavoring, browning, cooking, or preserving food, particularly meat, fish and tea, by exposing it to smoke from burning or smoldering material, most often wood. In Europe, alder is the traditional smoking wood, but ...

References

{{Taxonbar, from=Q2359163 Haemulinae Taxa named by Balthazar Osório Fish described in 1893