''Dactylosaurus'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

nothosaur Nothosaurs (superfamily Nothosauroidea) were Triassic marine sauropterygian reptiles. They averaged about in length, with a long body and tail. The feet were paddle-like, and are known to have been webbed in life, to help power the animal when sw ...

in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Pachypleurosauridae. Along with '' Anarosaurus'', ''Dactylosaurus'' was one of the earliest known pachypleurosaurs to come from

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

.Lepidosauromorpha: Pachypleurosauridae: Dactylosaurus & Anarosaurus
Palaeos.com. Last accessed 2008-07-04.

Etymology

''Dactylosaurus'' comes from the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

' (), "

finger A finger is a prominent digit (anatomy), digit on the forelimbs of most tetrapod vertebrate animals, especially those with prehensile extremities (i.e. hands) such as humans and other primates. Most tetrapods have five digits (dactyly, pentadact ...

" and ' (), meaning "

lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...

" or "

reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...

Description

''Dactylosaurus'' was a small reptile measuring up to long. The nasal bones of ''Dactylosaurus'' meet and are broadly structured.Rieppel, O & L Kebang (1995), "Pachypleurosaurs (Reptilia, Sauropterygia) from the Lower Muschelkalk, and a review of the Pachypleurosauroidea." ''Fieldiana Geol.'' N.S. 32: 1-44. The upper temporal fenestra is large and

kidney In humans, the kidneys are two reddish-brown bean-shaped blood-filtering organ (anatomy), organs that are a multilobar, multipapillary form of mammalian kidneys, usually without signs of external lobulation. They are located on the left and rig ...

-shaped. There are 17 cervical

vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spina ...

e and the cervical

rib In vertebrate anatomy, ribs () are the long curved bones which form the rib cage, part of the axial skeleton. In most tetrapods, ribs surround the thoracic cavity, enabling the lungs to expand and thus facilitate breathing by expanding the ...

s have anterior processes. The

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

e of ''Dactylosaurus'' extended broadly up the side of the snout.

''D. gracilis''

The

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

specimen ( MGUWR WR 3871s) of ''D. gracilis'' was only a partial

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

, consisting of the

anterior Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

end alone. Because it differed slightly from the fossils of ''D. gracilis,'' it was first thought to belong to the species ''D. schroederi'', which is now considered a

junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

for juvenile ''D. gracilis''. Once this was established, the juvenile fossil, which was found before the adult fossils, became the holotype. The one

limb Limb may refer to: Science and technology *Limb (anatomy), an appendage of a human or animal *Limb, a large or main branch of a tree *Limb, in astronomy, the curved edge of the apparent disk of a celestial body, e.g. lunar limb *Limb, in botany, t ...

that was found (a left forelimb), was noted to have a slimmer

radius In classical geometry, a radius (: radii or radiuses) of a circle or sphere is any of the line segments from its Centre (geometry), center to its perimeter, and in more modern usage, it is also their length. The radius of a regular polygon is th ...

and

ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...

than '' Neusticosaurus'', a similar nothosaur from

. ''D. gracilis'' is the smallest known species in its family, which includes the much more recognized '' Keichousaurus'', a nothosaur often remembered for its small size. The original holotype of ''D. gracilis'' is considered a juvenile, however the size of a nothosaur when its

bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...

s harden is used to show size, making the estimate as smallest member of its family still valid.

Distribution

''Dactylosaurus'' lived in the Early and Middle

Triassic period The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is the ...

during the Late

Olenekian In the geologic timescale, the Olenekian is an age (geology), age in the Early Triassic epoch (geology), epoch; in chronostratigraphy, it is a stage (stratigraphy), stage in the Lower Triassic series (stratigraphy), series. It spans the time betw ...

and

Anisian In the geologic timescale, the Anisian is the lower stage (stratigraphy), stage or earliest geologic age, age of the Middle Triassic series (stratigraphy), series or geologic epoch, epoch and lasted from million years ago until million years ag ...

faunal stage In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by conven ...

, of central Europe. In terms of

geology Geology (). is a branch of natural science concerned with the Earth and other astronomical objects, the rocks of which they are composed, and the processes by which they change over time. Modern geology significantly overlaps all other Earth ...

, they are found: 1) in the uppermost Röt (uppermost Buntsandstein; Lower Triassic): e.g. Michałkowice (Siemianowice Śląskie) and Kamień Śląski, S Poland, (the second location is not sure because Röt is not exposed there), 2) in the lowermost

Muschelkalk The Muschelkalk (German for "shell-bearing limestone"; ) is a sequence of sedimentary rock, sedimentary rock strata (a lithostratigraphy, lithostratigraphic unit) in the geology of central and western Europe. It has a Middle Triassic (240 to 230 m ...

(Middle Triassic), inter alia in the

Gogolin Formation Gogolin Formation – Triassic geologic formation, hitherto named the Gogolin Beds,Assmann P., 1913 – Beitrag zur Kenntniss der Stratigraphie des oberschlesischen Muschelkalks. Jb. Preuss. Geol. Landesanst., 34: 658 – 671, Berlin.Assmann P., ...

- Gogolin and its vicinity, S Poland.Sues H-D., Carroll R.L. 1985. The pachypleurosaurid Dactylosaurus schroederi (Diapsida: Sauropterygia). Canadian Journal of Earth Sciences 22(11): 1602-1608 In 2012, the new Röt site (~ 247 Ma; Lower Triassic; the latest Olenekian) with abundant disarticulated remains of ''Dactylosaurus'' was found in Gogolin. Presently, this in the only site where ''Dactylosaurus'' remains are accessible to collect.Kowal-Linka M., Bodzioch A. 2012a. Warstwy kościonośne z Dactylosaurus (Reptilia, Sauropterygia) z retu (trias dolny, olenek) Opolszczyzny ''Bonebeds with Dactylosaurus (Reptilia, Sauropterygia) from the Röt (Lower Triassic, Olenekian) in the Opole Silesia region (S Poland)''. Przegląd Geologiczny 60 (12): 646–649Kowal-Linka M., Bodzioch A. 2012b. New findings of vertebrate remains from the Röt (Lower Triassic, Olenekian) in the vicinity of Gogolin (Opole Silesia, S Poland). :Krasiejów - inspiracje paleontologiczne / Krasiejów - paleontological inspirations. E. Jagt-Yazykova, J. Jagt, A. Bodzioch, D. Konietzko-Meier (red.). Zakład Poligraficzno-Wydawniczy "Plik", Bytom: 70-80. Kowal-Linka M., Bodzioch A. 2017. Genesis of the Lower Triassic bonebeds from Gogolin (S Poland): The impact of microbial mats on trapping of vertebrate remains. Palaeogeography, Palaeoclimatology, Palaeoecology 466, 38–58

References

External links

Pachypleurosauridae
from Palaeos.com (technical)

from the Plesiosaur Directory * http://www.nrcresearchpress.com/doi/abs/10.1139/e85-169

Kowal-Linka & Bodzioch 2012b
{{Taxonbar, from=Q1391940 Early Triassic reptiles of Europe Triassic plesiosaurs Middle Triassic reptiles of Europe Anisian life Sauropterygian genera Fossil taxa described in 1884