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Cystiscidae is a
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. A ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of minute sea snails,
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
gastropod
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is es ...
s or
micromollusk A micromollusk is a shelled mollusk which is extremely small, even at full adult size. The word is usually, but not exclusively, applied to marine mollusks, although in addition, numerous species of land snails and freshwater mollusks also ...
s.


Description

The shell is minute to large, either white, uniformly colored, or patterned; the surface is smooth,
sculptured Sculptured is an American experimental and progressive metal band, mixing melodic and atonal segments. Biography Following the dissolution of his previous band, guitarist Don Anderson began Sculptured as mostly a one-man project in 1996.  With ...
, or axially costate; the
spire A spire is a tall, slender, pointed structure on top of a roof of a building or tower, especially at the summit of church steeples. A spire may have a square, circular, or polygonal plan, with a roughly conical or pyramidal shape. Spires are ...
is flat to immersed, or low to tall; the
protoconch A protoconch (meaning first or earliest or original shell) is an embryonic or larval shell which occurs in some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod. In older texts it is also calle ...
is paucispiral; the lip is thickened, smooth or denticulate; an external
varix A varix (pl. varices) is an abnormally dilated vessel with a tortuous course. Varices usually occur in the venous system, but may also occur in arterial or lymphatic vessels. Examples of varices include: * Varicose veins, large tortuous veins ...
is present or absent; a
siphonal notch A siphonal notch is a feature of the shell anatomy in some sea snails, marine gastropod mollusks. In these particular groups of sea snails the animal has a soft tubular anterior extension of the mantle called a siphon through which water is ...
is present or absent; a posterior notch is present or absent; the columella is multiplicate, internal
whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). Whorls in nature File:Photograph and axial plane floral d ...
cystiscid or modified cystiscid type. Mantle cavity with monopectinate ctenidium and bipectinate osphradium. Proboscis pleurembolic; jaws absent; typical
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
r sac present.


Taxonomy

The family was erected by
William Stimpson William Stimpson (February 14, 1832 – May 26, 1872) was a noted American scientist. He was interested particularly in marine biology. Stimpson became an important early contributor to the work of the Smithsonian Institution and later, dire ...
for a single included species based on unusual features of the head and
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
. Subsequent authors included this group in the family
Marginellidae Marginellidae, or the margin shells, are a taxonomic family of small, often colorful, sea snails, marine gastropod molluscs in the clade Neogastropoda. Taxonomy The higher classification of the family Marginellidae has long been in a state ...
. Coan (1965) recognized this group as a subfamily, but he did not include such genera as ''Gibberula'', ''Persicula'', and ''Canalispira''. Rule of priority of the
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the Int ...
require that the name Cystiscidae must be used, but this is unfortunate because the
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * Ty ...
specimen of the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is lost, and that species is poorly known.


2005 taxonomy

The family Cystiscidae consists of subfamilies (according to the Working Classification of the Gastropoda by Bouchet et al., 2005): * Canalispirinae Fedosov, Caballer & Bouchet, 2019 *
Cystiscinae Cystiscinae are a taxonomic subfamily of minute sea snails. These are marine gastropod mollusks or micromollusks in the family Cystiscidae, and the clade Neogastropoda. Diagnosis Shell minute to small, white; spire immersed to low; surface sm ...
Stimpson, 1865 * Granulininae G.A. & H.K. Coovert, 1995 *
Persiculinae ''Persiculinae'' is a taxonomic subfamily of minute to small predatory sea snails, marine gastropod mollusks. This subfamily includes several species which are micromollusks. Taxonomy The subfamily is sometimes placed in the family Cystiscidae, ...
G.A. & H.K. Coovert, 1995 *
Plesiocystiscinae Plesiocystiscinae is a subfamily of minute sea predatory sea snails, marine gastropod mollusks or micromollusks in the infraorder Neogastropoda. The genus name on which this subfamily name is based is derived from the Greekword ''plesios'', n ...
G.A. & H.K. Coovert, 1995Coovert G. A. & Coovert H. K. (1995). "Revision of the Supraspecific Classification of Marginelliform Gastropods". '' The Nautilus'' 109(2&3): 43-110, page 66, 70, 73. Reference for inclusion of Granulinae in Cystiscidae: Bouchet, P.; Rocroi, J.-P.; Fryda, J.; Hausdorf, B.; Ponder, W.; Valdes, A.; Waren, A. 2005. Classification and Nomenclator of Gastropod Families. Malacologia: International Journal of Malacology. Hackenheim, Germany: ConchBooks.


Genera

Genera within the family Cystiscidae include: Canalispirinae *''
Canalispira ''Canalispira'' is a genus of very small sea snails, marine gastropod mollusks or micromollusks in the family Cystiscidae. Species Species within the genus ''Canalispira'' include: *'' Canalispira attentia'' ( Laseron, 1957) *'' Canalispira aure ...
'' Jousseaume, 1875:168,270 *''
Osvaldoginella ''Osvaldoginella'' is a genus of sea snails, marine (ocean), marine gastropod mollusks in the family (biology), family Cystiscidae. Species Species within the genus ''Osvaldoginella'' include: * ''Osvaldoginella acualina'' Espinosa & Ortea, 201 ...
'' Espinosa & Ortea, 1997
Cystiscinae Cystiscinae are a taxonomic subfamily of minute sea snails. These are marine gastropod mollusks or micromollusks in the family Cystiscidae, and the clade Neogastropoda. Diagnosis Shell minute to small, white; spire immersed to low; surface sm ...
*
A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...
''
Topaginella ''Topaginella'' is a genus of minute fossil sea snails, marine gastropod mollusks or micromollusks in the family Cystiscidae. Like all of the related genera, this genus is sometimes still classified in the family Marginellidae. This genus ...
'' Laseron, 1957:288 *''
Crithe ''Crithe'' is a genus of minute sea snails, marine gastropod mollusks or micromollusks in the family Cystiscidae. This genus is sometimes still placed within the Marginellidae. These tiny gastropods have a shell which is colorless and is tran ...
''
Gould Gould may refer to: People * Gould (name), a surname Places United States * Gould, Arkansas, a city * Gould, Colorado, an unincorporated community * Gould, Ohio, an unincorporated community * Gould, Oklahoma, a town * Gould, West Virginia, ...
, 1860
*''
Cystiscus Cystiscus is a taxonomic genus of minute sea snails, marine gastropod mollusks or micromollusks. This genus was placed in the family Cystiscidae by Coovert and Coovert, 1995. Previously the genus was in the family Marginellidae, where it is sti ...
'' Stimpson, 1865:55 *''
Extra Extra or Xtra may refer to: Arts, entertainment and media Film * ''The Extra'' (1962 film), a Mexican film * ''The Extra'' (2005 film), an Australian film Literature * ''Extra'' (newspaper), a Brazilian newspaper * '' Extra!'', an American m ...
'' Jousseaume, 1894 - with the only species '' Extra extra'' Jousseaume, 1894 *'' Gibberula'' Swainson, 1840 *''
Inbiocystiscus ''Inbiocystiscus'' is a genus of minute sea snail, a marine gastropod mollusc in the family Cystiscidae Cystiscidae is a taxonomic family of minute sea snails, marine gastropod mollusks or micromollusks. Description The shell is minute to ...
'' Ortea and Espinosa, 2001 *''
Intelcystiscus ''Intelcystiscus'' is a genus of minute sea snail, a marine gastropod mollusc or micromollusc in the family Cystiscidae Cystiscidae is a taxonomic family of minute sea snails, marine gastropod mollusks or micromollusks. Description The she ...
'' Ortea and Espinosa, 2001 *
A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ...
''
Marginocystiscus ''Marginocystiscus'' is an extinct genus of sea snails, marine gastropod mollusks, in the family Cystiscidae.MolluscaBase (2018). ''Marginocystiscus'' Landau, da Silva & Heitz, 2016 †. Accessed through: World Register of Marine Species ...
'' Landau, C. M. Silva & Heitz, 2016 *''
Pachybathron ''Pachybathron'' is a genus of very small sea snails, marine gastropod mollusk or micromollusk in the subfamily Cystiscinae of the family Cystiscidae. Species Species within the genus ''Pachybathron'' include: *'' Pachybathron cassidiforme'' G ...
'' Gaskoin, 1853 *''
Persicula ''Persicula'' is a taxonomic genus of minute to small predatory sea snails, marine gastropod mollusks. It includes several species which are micromollusks.Persicula Schumacher, 1817. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine ...
''
Schumacher Schumacher or Schuhmacher is an occupational surname (German, "shoemaker", pronounced , both variants can be used as surnames, with Schumacher being the more popular one, however, only the variant with three "h"s can also be used as a job descript ...
, 1817
*''
Ticocystiscus ''Ticocystiscus'' is a genus of minute sea snail, a marine gastropod mollusc in the family Cystiscidae Cystiscidae is a taxonomic family of minute sea snails, marine gastropod mollusks or micromollusks. Description The shell is minute to ...
'' Espinosa and Ortea, 2002 *''
Ticofurcilla ''Ticofurcilla'' is a genus of very small sea snails, marine gastropod mollusks or micromollusks in the family Cystiscidae Cystiscidae is a taxonomic family of minute sea snails, marine gastropod mollusks or micromollusks. Description T ...
'' Espinosa & Ortea, 2002 ;Synonyms *''Furcilla'' Espinosa & Ortea, 2000: synonym of ''Ticofurcilla'' Espinosa & Ortea, 2002 (Invalid: junior homonym of '' Furcilla'' Martin, 1975 ipteraand Furcilla Bakharev, 1988 stracoda ''Ticofurcilla'' is a replacement name) **''Furcilla tica'' Espinosa & Ortea, 2000: synonym of ''
Ticofurcilla tica ''Ticofurcilla tica'' is a species of very small sea snail, a marine gastropod mollusk or micromollusk in the family Cystiscidae Cystiscidae is a taxonomic family of minute sea snails, marine gastropod mollusks or micromollusks. Descrip ...
'' (Espinosa & Ortea, 2000) Plesiocystiscinae * '' Plesiocystiscus'' Coovert & Coovert, 1995 Abbreviations: * (M) - original designation by monotype * OD - original designation * OD (M) - original designation as type species, but also monotypic * SD - subsequent designation as type * SD (M) - subsequent designation as type species, but also monotypic * T - original designation by tautonomy Comparisons of shells of the genera: File:Plesiocystiscus.jpg, File:Cystiscus.jpg, File:Crithe.jpg, File:ExtraExtra.jpg, File:Persicula.jpg, File:Gibberula.jpg, File:Canalispira.jpg, File:Furcilla.jpg, File:Granulina.jpg, File:Pugnus.jpg,


References

* Carpenter P. P. (1857) (1855-1857). ''Catalogue of the collection of Mazatlan shells in the British Museum collected by Frederick Reigen''. London:xvi+552 pp. (July 1855 - June 1857, each part dated at foot of first page) * Espinosa J & Ortea J. (2000). "Descripción de un género y once especies nuevas de Cystiscidae y Marginellidae (Mollusca: Neogastropoda) del Caribe de Costa Rica". ''Aviccenia'' (12/13): 95-114. * Espinosa J & Ortea J. (2002). "Nuevas especies de margineliformes de Cuba, Bahamas y el Mar Caribe de Costa Rica". ''Avicennia'' 15: 101-128. * Gould A. A. (1860). "Descriptions of shells collected in the North Pacific Exploring Expedition under Captains Ringold and Rogers. One part." ''
Proceedings Boston Society Natural History In academia and librarianship, conference proceedings is a collection of academic papers published in the context of an academic conference or workshop. Conference proceedings typically contain the contributions made by researchers at the confer ...
'' 7: 382-385. * Laseron C. F. (1957). "A new classification of the Australian Marginellidae (Mollusca), with a review of species from the Solanderian and Dampierian zoogeographical provinces". ''
Australian Journal of Marine and Freshwater Research Australian(s) may refer to: Australia * Australia, a country * Australians, citizens of the Commonwealth of Australia ** European Australians ** Anglo-Celtic Australians, Australians descended principally from British colonists ** Aboriginal Au ...
'' 8(3): 274-311. * Nevill G. & Nevill H. (1874). "Descriptions of new Mollusca from the Indian Ocean". ''
Journal Asiatic Society of Bengal A journal, from the Old French ''journal'' (meaning "daily"), may refer to: * Bullet journal, a method of personal organization * Diary, a record of what happened over the course of a day or other period *Daybook, also known as a general journal ...
'' Vol. XLIII, Part II. * Ortea J. & Espinosa J. (2001). "''Intelcystiscus'' e ''Inbiocystiscus'' (Mollusca: Neogastrópoda: Cystiscidae) dos nuevos géneros del Atlántico occidental tropical". ''Avicennia'' 14: 107-114. * Smiriglio C. & Mariottini P. (2001). "Description of ''Paolaura'' gen. nov. and of three new species of Cystiscidae from the Indian Ocean". ''
La Conchiglia ''La Conchiglia'' (1992), or ''The Conch'', is a short film from Somali writer and director Abdulkadir Ahmed Said. Plot A female artist is painting a seascape on a deserted beach when she discovers a beautiful conch. Curious, she puts her ear t ...
'' 299: 11-17. * Shumacher C. F. (1817). ''Essai d´un nouveau système des habitations des vers testacés''. Copenhagen, iv+287 pp., 22 plts. * Swainson W. (1840). ''A treatise on malacology, or the natural classification of shells and shellfish''. London, pp. 1–419, text figs. (Marginellidae: 323-324 in part 2)


External links


The Academy of Natural Sciences
A Database of Western Atlantic Marine Mollusca

{{Taxonbar, from=Q3009509 Gastropod families Taxa named by William Stimpson Volutoidea