The Cynipoidea are a moderate-sized
hymenoptera
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic.
Females typi ...
n superfamily that presently includes seven extant families and three extinct families, though others have been recognized in the past. The most familiar members of the group are
phytophagous, especially as
gall
Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants. Plant galls are abnormal outgrowths of plant tissues, similar to benign tumors or war ...
-formers, though the actual majority of included species are
parasitoid
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host (biology), host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionarily stable str ...
s or
hyperparasitoids. They are typically glossy, dark, smooth wasps with somewhat compressed bodies and somewhat reduced wing venation. It is common for various
metasoma
The metasoma is the posterior part of the body, or tagma (biology), tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory sy ...
l segments to be fused in various ways (often diagnostic for families or subfamilies), and the
petiole is very short, when present.
With the exception of the
Cynipidae (the gall wasps), it is a poorly known group as a whole, though there are nearly 3000 known species in total, and a great many species are still undescribed, mostly in the
Figitidae.
Each of the constituent families differs in biology, though life histories of one of the families (
Liopteridae) are still largely unknown. In July 2020, an identification key for the superfamily was published in the journal ''Insect Systematics and Diversity'', enabling identification to the family level.
Several groups formerly included in Cynipidae were elevated to family status in 2023.
Classification
::: Family
Protimaspidae Liu & Engel, 2007
::: Family
Stolamissidae Liu & Engel, 2007
::: Family
Gerocynipidae Liu & Engel, 2007
::: Family
Austrocynipidae
''Austrocynips mirabilis'' is a species of parasitic wasp. It is the only species in the genus ''Austrocynips'' and the family Austrocynipidae. It was originally described as a subfamily of Cynipidae; however, it was promoted to family status i ...
Riek, 1971
::: Family
Paraulacidae Nieves-Aldrey & Liljeblad, 2009
::: Family
Diplolepididae Latreille, 1802
::::: Subfamily
Diplolepidinae Latreille, 1802
::::: Subfamily
Pediaspinae Ashmead, 1903
::: Family
Ibaliidae Thomson, 1862
::::: Subfamily
Archaeibaliinae Liu & Engel, 2010
::::: Subfamily
Ibaliinae Thomson, 1862
::: Family
Liopteridae Ashmead, 1895
::::: Subfamily
Proliopterinae Liu & Engel, 2007
::::: Subfamily
Goeraniinae Liu & Engel, 2007
::::: Subfamily
Mayrellinae Hedicke, 1922
::::: Subfamily
Dallatorrellinae Kieffer, 1911
::::: Subfamily
Oberthuerellinae Hedicke, 1903
::::: Subfamily
Liopterinae Ashmead, 1895
::: Family
Figitidae Thomson, 1862
::::: Subfamily
Rasnicynipinae Kozlov, 1996
::::: Subfamily
Palaeocynipinae Kozlov, 1995
::::: Subfamily
Charipinae Dalla Torre & Kieffer, 1910
::::: Subfamily
Emargininae Kovalev, 1994
::::: Subfamily
Eucoilinae Thomson, 1862
:::::: Tribe
Diglyphosematini Belizin, 1961
:::::: Tribe
Eucoilini Thomson, 1862
:::::: Tribe
Ganaspini Belizin, 1961
:::::: Tribe
Kleidotomini Hellén, 1960
:::::: Tribe
Trichoplastini Kovalev, 1989
:::::: Tribe
Zaeucoilini Buffington, 2009
::::: Subfamily
Figitinae Thomson, 1862
::::: Subfamily
Mikeiinae Paretas-Martinez & Pujade-Villar, 2011
::::: Subfamily
Aspicerinae Dalla Torre & Kieffer, 1910
::::: Subfamily
Anacharitinae Thomson, 1862
::::: Subfamily
Pycnostigminae Cameron, 1905
::::: Subfamily
Thrasorinae Kovalev, 1994
::::: Subfamily
Plectocynipinae Ros-Farré & Pujade-Villar, 2007
::::: Subfamily
Euceroptresinae Buffington & Liljeblad, 2008
::::: Subfamily
Parnipinae Ronquist & Nieves-Aldrey, 2001
::: Family
Cynipidae Latreille, 1802
::::: Subfamily
Hodiernocynipinae Kovalev, 1994
::::: Subfamily
Cynipinae Latreille, 1802
:::::: Tribe
Aulacideini Nieves-Aldrey & Ronquist, 2015
:::::: Tribe
Aylacini Ashmead, 1903
:::::: Tribe
Ceroptresini Nieves-Aldrey, Nylander, & Ronquist, 2015
:::::: Tribe
Cynipini Latreille, 1802
:::::: Tribe
Diastrophini Nieves-Aldrey, Nylander, & Ronquist, 2015
:::::: Tribe
Eschatocerini Ashmead, 1903
:::::: Tribe
Phanacidini Nieves-Aldrey, Nylander, & Ronquist, 2015
:::::: Tribe
Qwaqwaiini Liljeblad, Nieves-Aldrey, & Melika, 2011
:::::: Tribe
Synergini Ashmead, 1896
References
Apocrita superfamilies
Hymenoptera superfamilies
{{Apocrita-stub