''Cyanothamnus quadrangulus'', commonly known as narrow-leaved boronia, is a plant in the citrus

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Rutaceae The Rutaceae () is a family (biology), family, commonly known as the rueRUTACEAE
in Bo ...

and is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to eastern

Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...

. It is an erect shrub with four-angled branches,

bipinnate The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade is divided into two or more leaflets ...

leaves and white, sometimes pale pink, four-

petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

led flowers. Boronia anethifolia habit

Description

''Cyanothamnus quadrangulus'' is an erect shrub that grows to a height of with four-angled,

glabrous Glabrousness () is the technical term for a lack of hair, down, setae, trichomes, or other such covering. A glabrous surface may be a natural characteristic of all or part of a plant or animal, or be due to loss because of a physical condition, ...

stems with prominent leaf scars. The leaves are bipinnate, long and wide in outline and have a

petiole Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, and ...

long. The leaves have between five and eleven glabrous, linear to narrow elliptic leaflets. The end leaflet is long and wide, the others usually slightly longer. The flowers are white, sometimes pale pink and are arranged in leaf

axil A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...

s, mainly in groups of between three and fourteen or more. The groups are borne on a peduncle long. The four

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s are triangular to broadly egg-shaped, about long and wide. The four

s are long with their bases overlapping. The eight

stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s have hairy edges. Flowering occurs from April to October and the fruit are glabrous, long and about wide.

Taxonomy and naming

This species was first formally described in 1863 by

Stephan Endlicher Stephan Friedrich Ladislaus Endlicher, also known as Endlicher István László (24 June 1804 – 28 March 1849), was an Austrian Empire, Austrian botanist, numismatist and Sinologist. He was a director of the Botanical Garden of Vienna. Biog ...

from an unpublished manuscript by Allan Cunningham and was given the name ''Boronia anethifolia''. Endlicher's description was published in ''

Enumeratio plantarum quas in Novae Hollandiae ora austro-occidentali ad fluvium Cygnorum et in sinu Regis Georgii collegit Carolus Liber Baro de Hügel ''Enumeratio plantarum quas in Novae Hollandiæ ora austro-occidentali ad fluvium Cygnorum et in sinu Regis Georgii collegit Carolus Liber Baro de Hügel'' is a description of the plants collected at the Swan River colony and King George Sound in ...

''. In a 2013 paper in the journal ''

Taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...

'',

Marco Duretto Marco Duretto (born 1964) is a manager and senior research scientist at the Royal Botanic Gardens Sydney in Australia. His primary research interests are systematics and conservation of Rutaceae, Rubiaceae, Orchidaceae, Stylidiaceae and evoluti ...

and others moved this species to the genus ''Cyanothamnus'' on the basis of

cladistic analysis Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...

, but because the name ''Cyanothamnus anethifolia'' had already been used for a different taxon (''Cyanothamnus anethifolius'' Bartl.) the name ''Cyanothamnus quadrangulus'' was used. The

specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

(''anethifolia'') referred to the deeply divided leaves with narrow leaflets which resemble those of

dill Dill (''Anethum graveolens'') is an annual herb in the celery family Apiaceae. It is native to North Africa, Iran, and the Arabian Peninsula; it is grown widely in Eurasia, where its leaves and seeds are used as a herb or spice for flavouring ...

(''Anethum'') in the

Apiaceae Apiaceae () or Umbelliferae is a family of mostly aromatic flowering plants named after the type genus ''Apium,'' and commonly known as the celery, carrot, or parsley family, or simply as umbellifers. It is the 16th-largest family of flowering p ...

and the new epithet (''quadrangulus'') means "four-angled" and refers to the four-sided stems of this species.

Distribution and habitat

This boronia grows in heath and forest and in rocky slopes and ridges between the Border Ranges in

Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...

and Wadbilliga Mountain in southern

New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...

References

{{Taxonbar, from=Q106902240 quadrangulus Flora of New South Wales Flora of Queensland Plants described in 2013