''Cyanea'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
s in the family
Campanulaceae
The family Campanulaceae (also bellflower family), of the order Asterales, contains nearly 2400 species in 84 genera of herbaceous plants, shrubs, and rarely small trees, often with milky sap. Among them are several familiar garden plants be ...
. The name ''Cyanea'' in
Hawaiian is ''hāhā''.
These
Hawaiian lobelioids
The Hawaiian lobelioids are a group of flowering plants in the bellflower family, Campanulaceae, subfamily Lobelioideae, all of which are endemic to the Hawaiian Islands. This is the largest plant radiation in the Hawaiian Islands, and indeed t ...
are
endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
to
Hawaii
Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only ...
with over 90% of ''Cyanea'' species are found only on one island in the Hawaiian chain.
[Givnish, T. J., et al. (2009)]
Origin, adaptive radiation and diversification of the Hawaiian lobeliads (Asterales: Campanulaceae).
''Proceedings of the Royal Society B: Biological Sciences'' 276(1656), 407-16. They grow in moist and wet forest habitat
[Lammers, T. G. (2004)]
Five new species of the endemic Hawaiian genus ''Cyanea'' (Campanulaceae: Lobelioideae).
''Novon'' 14(1) 84-101. and are largely
pollinated by birds
[ such as the ]Hawaiian honeycreeper
Hawaiian honeycreepers are a group of small, passerine birds endemic to Hawaii. They are closely related to the rosefinches in the genus ''Carpodacus'', but many species have evolved features unlike those present in any other finch. Their great ...
s,[ and the seeds are dispersed by birds that take the fruits.][ Most ''Cyanea'' are trees with few branches or none. The ]inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
is a raceme
A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the sh ...
of 4 to 45 flowers which grows from the leaf axils. The fruit is a fleshy berry
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, rasp ...
.[ There have been several theories regarding the evolution of large prickles on plants endemic to islands that lack any mammalian or reptilian herbivores. One such theory suggests that the prickles are a defense against herbivory by the ]moa-nalo
The moa-nalo are a group of extinct aberrant, goose-like ducks that lived on the larger Hawaiian Islands, except Hawaii itself, in the Pacific. They were the major herbivores on most of these islands until they became extinct after human settl ...
, a few taxa of flightless duck
Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a fo ...
s that went extinct on the islands within the last 1600 years.[Givnish, T. J., et al. (1994)]
Thorn-like prickles and heterophylly in ''Cyanea'': adaptations to extinct avian browsers on Hawaii?
''Proceedings of the National Academy of Sciences'' 91(7), 2810-14.
Species
There are about 78 species in the genus.[Oppenheimer, H. and D. H. Lorence. (2012)]
A new species of ''Cyanea'' (Campanulaceae, Lobelioideae) from Maui, Hawaiian Islands.
''PhytoKeys'' (13), 15-23. Species include:
* '' Cyanea aculeatiflora'' – Haleakala cyanea
* '' Cyanea acuminata'' – Honolulu cyanea
* '' Cyanea angustifolia'' – hāhā, aku,
* '' Cyanea arborea'' – palmtree cyanea
* ''Cyanea asarifolia
''Cyanea asarifolia'' is a rare species of flowering plant in the bellflower family known by the common name gingerleaf cyanea. It is endemic to Kauai, where there were no more than 30 individuals in a single population as of 2005.USFWS''Cyanea ...
'' – gingerleaf cyanea
* '' Cyanea aspleniifolia'' – spleenwort cyanea
* ''Cyanea calycina
''Cyanea calycina'' is a flowering plant in the Campanulaceae family. The IUCN has classified the species as critically endangered.
It is native to the Hawaiian Islands. An example is being monitored with a plant cam.
Description
It is a f ...
'' – Waianae Range rollandia, Oahu cyanea
* ''Cyanea comata
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Maui cyanea
* ''Cyanea copelandii
''Cyanea copelandii'' is a rare species of flowering plant in the bellflower family known by the common name treetrunk cyanea. It is endemic to Maui, where there are no more than 250 individuals remaining in the wild. It is a federally listed en ...
'' – treetrunk cyanea
* ''Cyanea coriacea
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – leatherleaf cyanea
* ''Cyanea crispa
''Cyanea crispa'' is a rare species of flowering plant known by the common names crimped rollandia and Koolau Range rollandia. It is endemic to Oahu, where there are no more than fifty individuals remaining in the Koʻolau Range.Bruegmann, M. M. ...
'' – Koolau Range rollandia
* ''Cyanea cylindrocalyx
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Splitleaf cyanea
* '' Cyanea dolichopoda'' – long-foot cyanea
* ''Cyanea dunbariae
''Cyanea dunbariae'' (often misspelled ''dunbarii'')USFWS''Cyanea dunbariae'' Five-year Review.January 2008. is a rare species of flowering plant in the bellflower family known by the common name ravine cyanea. It is endemic to Molokai, where th ...
'' – ravine cyanea
* '' Cyanea duvalliorum''
* ''Cyanea eleeleensis
''Cyanea eleeleensis'' was a rare species of flowering plant in the bellflower family known by the common name Eleele cyanea. It was endemic to Kauai, where it has been declared extinct.USFWSDetermination of endangered status for 48 species on ...
'' – eleele cyanea
* ''Cyanea elliptica
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – ellipticleaf cyanea
* ''Cyanea fissa
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Kauai cyanea
* '' Cyanea floribunda'' – Degener's cyanea
* ''Cyanea giffardii
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Kilauea Mauna cyanea
* '' Cyanea gibsonii''
* ''Cyanea glabra
''Cyanea glabra'' is a rare species of flowering plant in the family Campanulaceae known by the common name smooth cyanea. It is Endemism, endemic to Maui, where there are twelve plants remaining in the wild.Bruegmann, M. M. & V. Caraway. (2003)' ...
'' – smooth cyanea
* ''Cyanea grimesiana
''Cyanea grimesiana'' is a rare species of flowering plant in the bellflower family known by the common name splitleaf cyanea. It is native to Oahu and Molokai, where it is known from 12 occurrences.[Cyanea habenata
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...]
'' – stream-bed cyanea
* ''Cyanea hamatiflora
''Cyanea hamatiflora'' is a rare species of flowering plant in the bellflower family known by the common name wetforest cyanea. It is endemic to Hawaii, where it is known from the islands of Maui and Hawaii, and there are probably fewer than ...
'' – wetforest cyanea
* ''Cyanea hardyi
''Cyanea hardyi'', known in Hawaiian as hāhā, is a species of flowering plant in the bellflower family, Campanulaceae. This Hawaiian lobelioid is endemic to southern Kauai
Kauai, () anglicized as Kauai ( ), is geologically the second ...
'' – Oahu cyanea
* '' Cyanea heluensis''
* ''Cyanea hirtella
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – rustyleaf cyanea
* ''Cyanea horrida
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – prickly cyanea
* ''Cyanea humboldtiana
''Cyanea humboldtiana'' (formerly ''Rollandia humboldtiana'') is a rare species of flowering plant in the bellflower family known by the common name Oʻahu rollandia. It is native to Oʻahu, where it is known only from the Koʻolau Mountains. '' – Oahu rollandia
* '' Cyanea kahiliensis'' – spoonleaf cyanea
* '' Cyanea kauaulaensis''[
* '']Cyanea kolekoleensis
''Cyanea kolekoleensis'' (formerly ''Delissea kolekoleensis'') is a rare species of flowering plant in the bellflower family known by the common names ''`Oha'' and Kolekole cyanea. It is endemic to Kauai where it has only been seen in the Wahia ...
'' – kolekole cyanea
* ''Cyanea konahuanuiensis
''Cyanea konahuanuiensis'', known by the common name of Hāhā mili‘ohu (English: ''the ''Cyanea'' that is caressed by the mist''), is a species of plant from the Ko‘olau Mountains of O‘ahu, Hawaiian Islands. It was described in 2015, and ...
'' – Hāhā miliohu
* ''Cyanea koolauensis
''Cyanea koolauensis'' (formerly ''Rollandia angustifolia'') is a rare species of flowering plant in the bellflower family known by the common names Palolo Valley rollandia and narrowleaf rollandia. It is native to Oahu, where it is known only ...
'' – Palolo Valley rollandia
* ''Cyanea kuhihewa
''Cyanea kuhihewa'' is a rare species of flowering plant in the bellflower family known by the common name Limahuli Valley cyanea. It is endemic to Kauai, where only two mature plants are known from a single wild population. Like other '' Cyane ...
'' – Limahuli Valley cyanea
* ''Cyanea kunthiana
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Kunth's cyanea
* ''Cyanea lanceolata
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – lanceleaf cyanea
* ''Cyanea leptostegia
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – giant kokee cyanea
* '' Cyanea linearifolia'' – linearleaf cyanea
* '' Cyanea lobata'' – Waihee Valley cyanea
* ''Cyanea longiflora
''Cyanea longiflora'' (formerly ''Rollandia longiflora'') is a rare species of flowering plant in the bellflower family known by the common name ridge rollandia. It is endemic to Oahu where there are only three remaining occurrences in the nor ...
'' – ridge rollandia
* ''Cyanea longissima
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – streambank cyanea
* ''Cyanea macrostegia
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – purple cyanea
* ''Cyanea magnicalyx
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
''
* ''Cyanea mannii
''Cyanea mannii'' is a rare species of flowering plant in the bellflower family known by the common name Mann's cyanea. It is endemic to Hawaii, where it is known only from the island of Molokai. There are 9 occurrences with fewer than 3000 plan ...
'' – Mann's cyanea
* '' Cyanea maritae''
* ''Cyanea marksii
''Cyanea marksii'', or Marks' cyanea, is a tree in the bellflower family. It is critically endangered. Some think that it is extinct.
Distribution
It is native to North America and was found on Hawaii.
Taxonomy
It was named by Joseph Rock ...
'' – Marks' cyanea
* '' Cyanea mauiensis'' – Maui cyanea
* ''Cyanea mceldowneyi
''Cyanea mceldowneyi'' is a rare species of flowering plant in the Campanulaceae, bellflower family known by the common name McEldowney cyanea. It is Endemism, endemic to Hawaii, where it is known only from the island of Maui. '' – McEldowney's cyanea
* ''Cyanea membranacea
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – papery cyanea
* '' Cyanea minutiflora''
* '' Cyanea munroi'' – Munro's cyanea
* ''Cyanea obtusa
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – bluntlobe cyanea
* ''Cyanea parvifolia
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Waioli Valley rollandia
* '' Cyanea pilosa'' – hairy cyanea
* '' Cyanea pinnatifida'' – sharktail cyanea
* '' Cyanea platyphylla'' – puna cyanea
* ''Cyanea pohaku
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – pohaku cyanea
* ''Cyanea procera
''Cyanea procera'' is a rare species of flowering plant in the bellflower family known by the common name Molokai cyanea. It is endemic to Hawaii, where it is known only from the island of Molokai. It is a federally listed endangered species o ...
'' – Molokai cyanea
* ''Cyanea profuga
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Mapulehu Valley cyanea
* ''Cyanea pseudofauriei
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – hāhā
* ''Cyanea purpurellifolia
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – Panaluu Mountain rollandia
* '' Cyanea pycnocarpa'' – manyfruit cyanea
* ''Cyanea quercifolia
''Cyanea quercifolia'', known as oakleaf cyanea, was a species of plant native to the Hawaiian island of Maui. The plant is now considered extinct
Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a ...
'' – oakleaf cyanea
* ''Cyanea recta
''Cyanea recta'' is a rare species of flowering plant in the bellflower family known by the common names upright cyanea and Kealia cyanea. It is endemic to Hawaii, where it is known only from the island of Kauai.USFWS''Cyanea recta''.The Nature ...
'' – Kealia cyanea
* '' Cyanea remyi'' – Remy's cyanea
* ''Cyanea rivularis
''Cyanea rivularis'' (syn. '' Delissea rivularis'')USFWS''Delissea rivularis'' Five-year Review.August 2010. is a rare species of flowering plant in the bellflower family known by the common name plateau cyanea. It is endemic to Hawaii, where it ...
'' – plateau cyanea
* '' Cyanea salicina'' – willow cyanea
* ''Cyanea scabra
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – harsh cyanea
* ''Cyanea sessilifolia
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – sessileleaf cyanea
* ''Cyanea shipmanii
''Cyanea shipmanii'' is a rare species of flowering plant in the bellflower family known by the common name Shipman's cyanea. It is endemic to the island of Hawaii, where it is known only from the windward slopes of Mauna Loa and Mauna Kea. '' – Shipman's cyanea
* '' Cyanea solanacea'' – popolo
* ''Cyanea solenocalyx
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – pua kala, molokai
* ''Cyanea spathulata
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – spoonleaf cyanea
* ''Cyanea st.-johnii
''Cyanea st.-johnii'' (formerly ''Rollandia st.-johnii'') is a rare species of flowering plant in the bellflower family known by the common name St. John's rollandia. It is endemic to Oahu, where it is known only from the Koʻolau Mountains.USF ...
'' – St. John's rollandia
* ''Cyanea stictophylla
''Cyanea stictophylla'' is a rare species of flowering plant in the bellflower family known by the common name Kaiholena cyanea. It is endemic to the island of Hawaii, where it is known only from the rainforests of Mauna Loa.World Conservation ...
'' – Kaiholena cyanea
* ''Cyanea superba
''Cyanea superba'' is a rare species of flowering plant in the bellflower family known by the common names Mt. Kaala cyanea and superb cyanea. It is endemic to the island of Oahu, but it is now extinct in the wild. It exists in cultivation and s ...
'' – Mt. Kaala cyanea
* ''Cyanea tritomantha
Cyanea may refer to:
* ''Cyanea'' (jellyfish), a genus of jellyfish in the family Cyaneidae
* ''Cyanea'' (plant), a genus of Hawaiian plants in the family Campanulaceae
* An ancient Greek name that means "blue sky" and in the Greek alphabet it' ...
'' – aku aku, aku
* ''Cyanea truncata
''Cyanea truncata'' is a rare species of flowering plant in the bellflower family known by the common name Punaluu cyanea. It is endemic to the islands of Oahu and Molokai in Hawaii, but it is now critically endangered. It exists in cultivation ...
'' – Punaluu cyanea
* ''Cyanea undulata
''Cyanea undulata'' is a rare species of flowering plant in the bellflower family known by the common names wavy cyanea and leechleaf cyanea. It is endemic to the island of Kauai, where it is in rapid decline.USFWS''Cyanea undulata'' Five-year R ...
'' – leechleaf cyanea
References
{{Taxonbar, from=Q148336
Endemic flora of Hawaii
Campanulaceae genera