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''Cupha erymanthis'', the rustic, is a
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of brush-footed
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
found in forested areas of tropical
South Asia South Asia is the southern subregion of Asia, which is defined in both geographical and ethno-cultural terms. The region consists of the countries of Afghanistan, Bangladesh, Bhutan, India, Maldives, Nepal, Pakistan, and Sri Lanka.;;;;; ...
and
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. The males and females are identical.


Description


Upperside

The upperside of the rustic is ochraceous light brown. Its forewing displays some loop-like, slender, dark cellular markings with a broad, somewhat curved, transverse yellow discal band from costa to vein 1. The band does not reach the termen but broadens posteriorly. The margins of the forewing are irregularly sinuous, with the inner defined broadly with black, and produced outwards in interspaces 3 and 4. Below this, the margin is squarely indented inwards in interspace 2 and outwardly convex in interspace 1. There is a curved series of three black spots. The largest is in interspaces 1, 2 and 3. The apical area beyond the band is black, with a conspicuous yellow subapical spot in interspace 5, and a paler ill-defined similar spot above it in interspace 6. In the posterior, the black area is produced narrowly to the tornus and encircles a yellow spot near the apex of interspace 2. The hindwing features a transverse sinuous and very slender black line. This line is followed by a slender and somewhat lunular line; a transverse discal series of five black spots in interspaces 2 to 6; a postdiscal medially disjointed series or broad black lunules; a subterminal series of similar but straighter lunges; and a narrow terminal black band. The outer subbasal transverse line broadens at the costa, and is outwardly margined by pale spots in the interspaces. These are anteriorly white and well defined, and posteriorly obscure or often absent.


Underside

The underside of the wing is much paler. The discal band on the forewing is pale, and the black lunules on the apical area are replaced by pale brownish ochre. There are some obscure markings in the cells of both the forewings and hindwings. Markings on the forewing include a discal, transverse, slender, chestnut-brown lunular line, bent inwards above vein 5, and bordered outwardly by a series of dark spots. The large black spot in interspace 1 is the same as that on the upperside. There are inner and outer transverse subterminal series of small dentate spots. On the hindwing are indistinct cellular markings. The outer subbasal dark transverse line is similar to that on the upperside, but is more clearly defined and very sinuous. There is a transverse discal series of uneven lunules, paler than the ground colour, followed by a series of dark spots. There is a postdiscal very obscure pale lunular band, and a subterminal series of dentate dark spots, often obscure or obsolescent. The antennae, head, thorax and abdomen are ochraceous brown. Beneath, the palpi, thorax, and abdomen are a very pale ochraceous white. The
caterpillar Caterpillars ( ) are the larva, larval stage of members of the order Lepidoptera (the insect order comprising butterfly, butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawfly ...
is brown, with a dorsal and lateral series of darker brown markings. The head has two slender branched spines. Succeeding segments on either side feature a lateral series of semitransparent similar brown spines. The
pupa A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their ...
is green, studded with eight slender pink filaments and four small pink tubercles.


Subspecies

* ''Cupha erymanthis erymanthis'' - China, Hong Kong, Taiwan * ''Cupha erymanthis placida'' Moore,
881 __NOTOC__ Year 881 ( DCCCLXXXI) was a common year starting on Sunday of the Julian calendar. Events By place Europe * February 12 – King Charles the Fat, the third son of the late Louis the German, is crowned as Holy Roman Emper ...
/small> - Sri Lanka * ''Cupha erymanthis maja'', Fruhstorfer (1898) – south India * ''Cupha erymanthis lotis'', Sulzer (1776) – northeast India, Thailand, Malaya, Singapore, Indochina * ''Cupha erymanthis andamanica'', Moore (1900) – Andamans * ''Cupha erymanthis nicobarica'', Felder (1862) – Nicobar Is.


Ecology

The rustic caterpillars feed mainly on
Flacourtiaceae The Flacourtiaceae is a defunct family of flowering plants whose former members have been scattered to various families, mostly to the Achariaceae and Salicaceae. It was so vaguely defined that hardly anything seemed out of place there and it bec ...
species, for example '' Flacourtia montana'', '' F. ramontchii'', '' F. rukam'', ''
Xylosma racemosa ''Xylosma'' is a genus of flowering plants in the family Salicaceae. It contains around 100 species of evergreen shrubs and trees commonly known as brushhollies, xylosmas, or, more ambiguously, "logwoods". The generic name is derived from the ...
'' and '' Scolopia'' species. They also eat plants such as ''
Glochidion eriocarpum ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the P ...
'' and '' Lepisanthes rubiginosum''. Adult butterflies occasionally visit
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
to drink the fluids. They seem to favour carcasses lying in exposed, sunny areas over those which are in the shade.Hamer, K. et al "Diversity and ecology of carrion and fruit feeding butterflies in Bornean rain forest". ''Journal of Tropical Ecology'' 2006 (22) p 25–33. DOI10.1017/S0266467405002750.


Larval Host Plants

* ''
Glochidion eriocarpum ''Glochidion'' is a genus of flowering plants, of the family Phyllanthaceae, known as cheese trees or buttonwood in Australia, and leafflower trees in the scientific literature. It comprises about 300 species, distributed from Madagascar to the P ...
'' * ''
Flacourtia ''Flacourtia'' is a genus of flowering plants in the family Salicaceae. It was previously placed in the now defunct family Flacourtiaceae. The generic name honors Étienne de Flacourt (1607–1660), a governor of Madagascar. It contains 15 s ...
'' * ''
Flacourtia indica ''Flacourtia indica'' ( syn. ''Flacourtia ramontchi''), known commonly as ramontchi, governor's plum, Madagascar plum and Indian plum, is a species of flowering plant native to much of Africa and tropical and temperate parts of Asia. ''F. indica' ...
'' * '' Flacourtia jangomas'' * '' Flacourtia montana''


Life history

File:Cupha erymanthis - Southern Rustic Butterfly caterpillar on Flacourtia montana.JPG, Caterpillar File:Cupha erymanthis – Sahyadri Rustic pupa 08.jpg, Pupa File:Cupha erymanthis-Kadavoor-2016-03-31-001.jpg, Pupa (matured) Cupha erymanthis at Kadavoor.jpg, ''C. e. maya'',
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South C ...
, India File:Southern Rustic @ Kanjirappally.jpg


References

{{Taxonbar, from=Q714578 Vagrantini Butterflies of Asia Butterflies of Indochina Butterflies of Singapore Taxa named by Dru Drury Butterflies described in 1773