The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...

, of which the most important to humans are: *'' Cucurbita'' – squash,

pumpkin A pumpkin is a vernacular term for mature winter squash of species and varieties in the genus ''Cucurbita'' that has culinary and cultural significance but no agreed upon botanical or scientific meaning. The term ''pumpkin'' is sometimes use ...

, zucchini, some gourds *'' Lagenaria'' – calabash, and others that are inedible *'' Citrullus'' – watermelon (''C. lanatus'', ''C. colocynthis'') and others *'' Cucumis'' –

cucumber Cucumber (''Cucumis sativus'') is a widely-cultivated Vine#Horticultural climbing plants, creeping vine plant in the Cucurbitaceae family that bears usually cylindrical Fruit, fruits, which are used as culinary vegetables.

(''C. sativus''), various melons and vines *'' Momordica'' – bitter melon *'' Luffa'' – the common name is also luffa, sometimes spelled loofah (when fully ripened, two species of this fibrous fruit are the source of the loofah scrubbing sponge) *'' Cyclanthera'' –

Caigua ''Cyclanthera pedata'', known as ''caigua'', is a herbaceous vine grown for its edible fruit, which is predominantly used as a vegetable. It is known from cultivation only, and its use goes back many centuries as evidenced by ancient phytomorph ...

The plants in this family are grown around the tropics and in temperate areas, where those with edible fruits were among the earliest cultivated plants in both the Old and New Worlds. The family Cucurbitaceae ranks among the highest of plant families for number and percentage of species used as human food. The name ''Cucurbitaceae'' comes to

international scientific vocabulary International scientific vocabulary (ISV) comprises scientific and specialized words whose language of origin may or may not be certain, but which are in current use in several modern languages (that is, translingually, whether in naturalized, loa ...

from New Latin, from '' Cucurbita'', the type genus, + '' -aceae'', a standardized

suffix In linguistics, a suffix is an affix which is placed after the stem of a word. Common examples are case endings, which indicate the grammatical case of nouns, adjectives, and verb endings, which form the conjugation of verbs. Suffixes can carry ...

for plant family names in modern taxonomy. The genus name comes from the Classical Latin word '' cucurbita'', "gourd".

Description

Most of the plants in this family are annual vines, but some are woody

liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...

s, thorny shrubs, or trees ('' Dendrosicyos''). Many species have large, yellow or white flowers. The stems are hairy and pentangular. Tendrils are present at 90° to the leaf petioles at nodes. Leaves are exstipulate, alternate, simple

palmately The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...

lobed or palmately compound. The flowers are unisexual, with male and female flowers on different plants (

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...

) or on the same plant (

monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...

). The female flowers have inferior ovaries. The fruit is often a kind of modified

berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, raspb ...

called a

pepo PEPO Lappeenranta (abbreviated PEPO) is a football club from Lappeenranta in Finland. The club was formed in 1958 and their home ground is at the Kimpinen Sports Centre. The men's first team currently plays in the Ykkönen (Second Division). ...

Fossil history

One of the oldest fossil cucurbits so far is †''Cucurbitaciphyllum lobatum'' from the Paleocene epoch, found at Shirley Canal, Montana. It was described for the first time in 1924 by the paleobotanist

Frank Hall Knowlton Frank Hall Knowlton (September 2, 1860 - November 22, 1926) was an American botanist, ornithologist and naturalist. Born in Vermont, he joined the Geological Survey and took an interest in fossil plants in the local lignite, later becoming a sp ...

. The fossil leaf is

palmate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...

, trilobed with rounded lobal sinuses and an entire or serrate margin. It has a leaf pattern similar to the members of the genera ''

Kedrostis ''Kedrostis'' is a genus of ± 35 species climbing or trailing herbs in the family Cucurbitaceae. Its native range is tropical Africa and Asia. Species # '' Kedrostis abdallai'' A. Zimmermann # ''Kedrostis africana'' (L.) Cogn. # ''Kedrosti ...

'', ''

Melothria ''Melothria'' is a genus of flowering plants in the family Cucurbitaceae, native to the Americas from the United States to Argentina, and with some introductions in Africa and elsewhere. A number of Old World species formerly in ''Melothria'' wer ...

'' and '' Zehneria''.

Classification

Tribal classification

2016-11-10 Cucurbits (Cucurbitaceae) Madrid Botanico

The most recent classification of Cucurbitaceae delineates 15 tribes: * Tribe Gomphogyneae Benth. & Hook.f. ** ''

Alsomitra ''Alsomitra'' is a genus of the plant family Cucurbitaceae (the cucumber, squash, and pumpkin family). The seeds of ''Alsomitra macrocarpa'' are among the largest winged seeds in the plant kingdom and their shape inspired a number of aviation pi ...

'' (Blume) Spach (1 sp.) ** '' Bayabusua'' (1 sp.) ** ''

Gomphogyne ''Gomphogyne'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the T ...

'' Griff. (2 spp.) ** ''

Gynostemma ''Gynostemma'' is a genus of perennial climbing vines in the Cucurbitaceae, cucumber, gourd, and melon family, comprising at least 19 species, all native to the tropical East or Far East, inclusive of the Himalayas: China (with 9 endemism, endemi ...

'' Blume (10 spp.) ** ''

Hemsleya ''Hemsleya'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Eastern Himalayas to southern China and Papuasia. It is native to the countries of Assam (in India), Bismarck Archipelago (island group, part ...

'' Cogn. ex F.B.Forbes & Hemsl. (30 spp.) ** '' Neoalsomitra'' Hutch. (12 spp.) * Tribe Triceratieae A.Rich. ** '' Anisosperma'' Silva Manso (1 sp.) ** ''

Cyclantheropsis ''Cyclantheropsis'' is a genus of flowering plants belonging to the family Cucurbitaceae The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95 genera, of which the most i ...

'' Harms (3 spp.) ** ''

Fevillea ''Fevillea'' is a genus of the plant family Cucurbitaceae (the cucumber, squash, and pumpkin family). Some references suggest that there is only one member of this genus, '' Fevillea cordifolia'' (antidote vine). Other online sources make refere ...

'' L. (8 spp.) ** ''

Pteropepon ''Pteropepon'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Western South America to Northwestern Argentina and Brazil. Species: *'' Pteropepon acaranthus'' *'' Pteropepon deltoideus'' *'' Pterop ...

'' (Cogn.) Cogn. (5 spp.) ** ''

Sicydium ''Sicydium'' is a genus of gobies native to fast-flowing streams and rivers of the Americas (Central America, Mexico, Cocos Island, the Caribbean, Colombia, Ecuador and Venezuela) with a couple species native to Middle Africa. Species There are ...

'' Schltdl. (7 spp.) * Tribe Zanonieae Benth. & Hook.f. ** ''

Gerrardanthus ''Gerrardanthus'' is a genus of flowering plants native to tropical Africa and South Africa, first described by William Henry Harvey (1811–1866), and named in honor of William Tyrer Gerrard (died 1866 in Mahavelona, Madagascar), botanical col ...

'' Harvey in Hook.f. (3–5 spp.) ** ''

Siolmatra ''Siolmatra'' is a genus of flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel' ...

'' Baill. (1 sp.) ** ''

Xerosicyos ''Xerosicyos'' is a flowering plant genus of the family (biology), family Cucurbitaceae. Its name comes from Greek language, Greek ''xeros'' (meaning "dry") and ''sicyos'' ("cucumber"). There are three species, all endemic to Madagascar. ''Xero ...

'' Humbert (5 spp.) ** ''

Zanonia ''Zanonia'' is a monotypic genus in the flowering plant family Cucurbitaceae (the cucumber, squash, and pumpkin family). The only species is ''Zanonia indica'', a medium-sized liana found in the Indian Subcontinent and Southeast Asia east to Ne ...

'' L. (1 sp.) * Tribe Actinostemmateae H.Schaef. & S.S.Renner ** ''

Actinostemma ''Actinostemma'' is a genus of flowering plants in the gourd family Cucurbitaceae, native to east Asia. They are slender, weakly twining/climbing annual herbs. Species Currently accepted species include: *'' Actinostemma lobatum'' (Maxim.) Max ...

'' Griff. (3 spp.) * Tribe Indofevilleeae H.Schaef. & S.S.Renner ** ''

Indofevillea ''Indofevillea'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Eastern Himalayas, Assam, to Tibet and Myanmar. The genus name of ''Indofevillea'' is in honour of Louis Feuillée (1660–1732), a French ...

Chatterjee Chatterjee or Chattopadhyay (চট্টোপাধ্যায়) is a Bengali Hindu family name, used primarily by the Kulin group of Pancha-Gauda Brahmins in India, and associated with the Bengali Brahmin caste. ''Chatterjee'' is an Anglicize ...

(2 sp.) * Tribe Thladiantheae H.Schaef. & S.S.Renner ** ''

Baijiania ''Baijiania'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Borneo. Species: *''Baijiania borneensis ''Baijiania'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native ...

'' A.M.Lu & J.Q.Li (30 spp.) ** '' Thladiantha'' Bunge 1833 (5 spp.) * Tribe Siraitieae H. Schaef. & S.S. Renner ** '' Siraitia'' Merr. (3–4 spp.) * Tribe Momordiceae H.Schaef. & S.S.Renner ** '' Momordica'' L. (60 spp.) * Tribe Joliffieae Schrad. ** '' Ampelosicyos'' Thouars (5 spp.) ** ''

Cogniauxia ''Cogniauxia'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Western Central Tropical Africa to Angola. Species: *''Cogniauxia podolaena'' *''Cogniauxia trilobata ''Cogniauxia'' is a genus of flo ...

'' Baill. (2 spp.) ** ''

Telfairia ''Telfairia'' is a small genus of flowering plants in the squash family which are native to Africa. They include ''Telfairia occidentalis'', the fluted pumpkin, which is an important vegetable in Nigeria and other African nations. It is also kno ...

'' Hook. (3 spp.) * Tribe Bryonieae Dumort. ** ''

Austrobryonia ''Austrobryonia'' is a genus of flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, ve ...

'' H.Schaef. (4 spp.) ** '' Bryonia'' L. (10 spp.) ** ''

Ecballium ''Ecballium'' is a genus of flowering plants in the family Cucurbitaceae containing a single species, ''Ecballium elaterium'', also called the squirting cucumber or exploding cucumber (not the same plant as ''Cyclanthera brachystachya''). Its u ...

'' A.Rich. (1 sp.) * Tribe Schizopeponeae C.Jeffrey ** ''

Herpetospermum ''Herpetospermum'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Himalaya The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from th ...

'' Wall. ex Hook.f. (3 spp.) ** '' Schizopepon'' Maxim. (6–8 spp.) * Tribe Sicyoeae Schrad. ** '' Cyclanthera'' Schrad. (40 spp.) ** '' Echinocystis'' Torr. & A.Gray (1 sp.) ** '' Echinopepon'' Naudin (20 spp., including '' Brandegea'' Cogn.) ** ''

Frantzia ''Frantzia'' is a genus of the gourd family, Cucurbitaceae containing around 6 species of herbaceous climbers with tuberous roots. All members grow in forest and secondary scrub in Central America where tacaco ('' Frantzia tacaco'') is a widely c ...

'' Pittier (5 spp.) ** '' Hanburia'' Seem. (7 spp.) ** '' Hodgsonia'' Hook.f. & Thomson (2 spp.) ** ''

Linnaeosicyos ''Linnaeosicyos'' is a monotypic genus of flowering plants belonging to the family Cucurbitaceae. The only species is ''Linnaeosicyos amara'' . It is native to the Dominican Republic. The genus name of ''Linnaeosicyos'' is in honour of Carl Linn ...

'' H.Schaef. & Kocyan (1 sp.) ** '' Luffa'' Mill. (5–7 spp.) ** '' Marah'' Kellogg (7 spp.) ** ''

Nothoalsomitra ''Nothoalsomitra suberosa'' is a species of subtropical vines in the family Cucurbitaceae, the only member of the monotypic genus ''Nothoalsomitra''. Native to Queensland in Australia. The species is a vigorous climber with pale corky bark and ...

'' Hutch. (1 sp.) ** '' Sicyos'' L. (75 spp., including '' Sechium'' P.Browne) ** ''

Trichosanthes ''Trichosanthes'' is a genus of tropical and subtropical vines. They belong to the cucumber family (Cucurbitaceae), and are closely related to '' Gymnopetalum''. ''Hodgsonia'', formerly included here, is usually considered a well-distinct genus ...

'' L. (≤100 spp.) * Tribe Coniandreae Endl. ** ''

Apodanthera ''Apodanthera'' is a genus of plants in the family Cucurbitaceae The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95 genera, of which the most important to humans are: ...

'' Arn. (16 spp.) ** '' Bambekea'' Cogn. (1 sp.) ** ''

Ceratosanthes ''Ceratosanthes'' is a genus of flowering plants belonging to the family Cucurbitaceae The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95 genera, of which the most imp ...

'' Adans. (4 spp.) ** '' Corallocarpus'' Welw. ex Benth. & Hook.f. (17 spp.) ** '' Cucurbitella'' Walp. (1 sp.) ** '' Dendrosicyos'' Balf.f. (1 sp.) ** '' Doyerea'' Grosourdy (1 sp.) ** '' Eureiandra'' Hook.f. (8 spp.) ** '' Gurania'' (Schltdl.) Cogn. (37 spp.) ** '' Halosicyos'' Mart.Crov (1 sp.) ** ''

Helmontia ''Helmontia'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is from northern South America to northern Brazil. It is found in the countries of Brazil, French Guiana, Guyana, Suriname and Venezuela. The ge ...

'' Cogn. (2–4 spp.) ** ''

Ibervillea ''Ibervillea'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Arizona, Oklahoma New Mexico, Texas and south to Mexico, Belize and Guatemala. The genus name of ''Ibervillea'' is in honour of Pierre Le Mo ...

'' Greene (9–10 spp.) ** ''

'' Medik. (28 spp.) ** '' Melotrianthus'' M.Crovetto (1–3 spp.) ** '' Psiguria'' Neck. ex Arn. (6–12 spp.) ** '' Seyrigia'' Keraudren (6 spp.) ** '' Trochomeriopsis'' Cogn. (1 sp.) ** '' Tumamoca'' Rose (2 spp.) ** '' Wilbrandia'' Silva Manso (5 spp.) * Tribe

Benincaseae Benincaseae is one of 15 Tribe (biology), tribes in the family Cucurbitaceae. Subtribes and genera Benincaseae consists of at least one subtribe, 26 genera, and over 200 species. *Benincasinae **''Acanthosicyos'' **''Benincasa'' **''Citrullus'' ...

Ser. ** ''

Acanthosicyos ''Acanthosicyos'' is a genus of thorny shrubs of the botanical family Cucurbitaceae, subfamily Cucurbitoideae. The genus name derives from the Greek words "''akantha''" for thorn and "''sykios''" for cucumber or gourd. Species Endemic to the Na ...

'' Welw. ex Hook.f. (1 sp.) ** '' Benincasa'' Savi (2 spp., including '' Praecitrullus'' Pangalo) ** '' Borneosicyos'' (1–2 spp.) ** '' Cephalopentandra'' Chiov. (1 sp.) ** '' Citrullus'' Schrad. (4 spp.) ** ''

Coccinia The scarlet gourds are a genus (''Coccinia'' from the Greek, ''kokkinia'' or ''kokkinias'' - "red" or "scarlet") with 25 species. It is distributed in sub- Saharan Africa and with one species, ''C. grandis'' also in South and South-east Asia, a ...

'' Wight & Arn. (30 spp.) ** '' Ctenolepis'' Hook. f. 1867 (3 spp.) ** '' Cucumis'' L. (65 spp.) ** ''

Dactyliandra ''Dactyliandra'' is a genus of flowering plants belonging to the family Cucurbitaceae The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95 genera, of which the most impo ...

'' Hook.f. (2 spp.) ** '' Diplocyclos'' (Endl.) T.Post & Kuntze (4 spp.) ** '' Indomelothria'' (2 spp.) ** ''

Khmeriosicyos ''Khmeriosicyos'' is a genus of flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, ves ...

'' (1 sp.) ** '' Lagenaria'' Ser. (6 spp.) ** '' Lemurosicyos'' Keraudren (1 sp.) ** ''

'' L. (12 spp., including '' M. scabra'') ** '' Muellerargia'' Cogn. (2 sp.) ** '' Papuasicyos'' (8 spp.) ** ''

Peponium ''Peponium'' is a genus of flowering plants belonging to the family Cucurbitaceae. Its native range is Tropical and Southern Africa, and Western Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, coveri ...

'' Engl. (20 spp.) ** '' Raphidiocystis'' Hook.f. (5 spp.) ** ''

Ruthalicia ''Ruthalicia'' is a genus of flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel ...

'' C.Jeffrey (2 spp.) ** '' Scopellaria'' W.J.de Wilde & Duyfjes (2 spp.) ** '' Solena'' Lour. (3 spp.) ** '' Trochomeria'' Hook.f. (8 spp.) ** '' Zehneria'' Endl. (''ca.'' 60 spp.) * Tribe Cucurbiteae Ser. ** ''

Abobra ''Abobra'' is a monotypic genus of the gourd family containing the one species ''Abobra tenuifolia'' ( syn. ''Abobra viridiflora'' Naudin, ''Bryonia tenuifolia'' Hook. & Arn.). It is a diecious, perennial climbing plant reaching up to 4 meters h ...

'' Naudin (1 sp.) ** '' Calycophysum'' H.Karst. & Triana (5 spp.) ** '' Cayaponia'' Silva Manso (50–59 spp., including '' Selysia'' Cogn.) ** '' Cionosicys'' Griseb. (4–5 spp.) ** '' Cucurbita'' L. (15 spp.) ** '' Penelopeia'' Urb. (2 spp.) ** ''

Peponopsis ''Peponopsis'' is a monotypic genus of flowering plants belonging to the family Cucurbitaceae The Cucurbitaceae, also called cucurbits or the gourd family, are a plant family consisting of about 965 species in around 95 genera, of which the mo ...

'' Naudin (1 sp.) ** '' Polyclathra'' Bertol. (6 spp.) ** '' Schizocarpum'' Schrad. (11 spp.) ** '' Sicana'' Naudin (4 spp.) ** '' Tecunumania'' Standl. & Steyerm. (1 sp.)

Systematics

Modern molecular phylogenetics suggest the following relationships: {, class="collapsible collapsed" style="width:100%; border:solid 1px #aaa" , - ! style="background:#F0F2F5" , Detailed Cladogram showing Cucurbitaceae phylogeny , - , {{Clade, style=line-height:90%;font-size:90%; , label1=Cucurbitaceae , 1={{clade , 1={{clade , label1= Gomphogyneae , 1={{clade , 1=''

'' , 2={{clade , 1='' Bayabusua'' , 2={{clade , 1={{clade , 1=''

'' , 2=''

'' , 2={{clade , 1=''

'' , 2='' Neoalsomitra'' , 2={{clade , 1={{clade , label1= Triceratieae , 1={{clade , 1=''

'' , 2={{clade , 1=''

'' , 2=''

'' , label2= Zanonieae , 2={{clade , 1=''

'' , 2={{clade , 1={{clade , 1=''

'' , 2=''

'' , 2={{clade , 1={{clade , label1= Actinostemmateae , 1=''

'' , 2={{clade , 1={{clade , label1= Indofevilleeae , 1=''

'' , 2={{clade , 1={{clade , label1= Thladiantheae , 1={{clade , 1=''

'' , 2='' Thladiantha'' , 2={{clade , 1={{clade , label1= Siraitieae , 1='' Siraitia'' , 2={{clade , 1={{clade , label1= Momordiceae , 1='' Momordica'' , 2={{clade , 1={{clade , label1= Joliffieae , 1={{clade , 1=''

'' , 2={{clade , 1=''

'' , 2='' Ampelosicyos'' , 2={{clade , 1={{clade , label1= Bryonieae , 1={{clade , 1={{clade , 1=''

'' , 2='' Bryonia'' , 2=''

'' , 2={{clade , 1={{clade , label1= Sicyoeae , 1={{clade , 1={{clade , 1=''

'' , 2={{clade , 1='' Hodgsonia'' , 2={{clade , 1={{clade , 1='' Echinocystis'' , 2='' Marah'' , 2={{clade , 1={{clade , 1='' Echinopepon'' , 2=''

'' , 2={{clade , 1='' Cyclanthera'' , 2='' Hanburia'' , 3='' Sicyos'' , 2={{clade , 1=''

'' , 2={{clade , 1='' Luffa'' , 2=''

'' , 2={{clade , label1= Schizopeponeae , 1={{clade , 1='' Schizopepon'' , 2=''

'' , 3={{clade , 1={{clade , label1= Coniandreae , 1=CONIANDREAE , 2={{clade , label1= Cucurbiteae , 1=CUCURBITEAE , label2=

, 2=BENINCASEAE , targetA=CONIANDREAE , subcladeA={{clade , 1={{clade , 1='' Bambekea'' , 2='' Eureiandra'' , 2={{clade , 1='' Dendrosicyos'' , 2={{clade , 1={{clade , 1='' Trochomeriopsis'' , 2='' Seyrigia'' , 2={{clade , 1='' Corallocarpus'' , 2={{clade , 1='' Cucurbitella'' , 2={{clade , 1={{clade , 1='' Doyerea'' , 2={{clade , 1='' Wilbrandia'' , 2={{clade , 1='' Psiguria'' , 2={{clade , 1=''

'' , 2='' Gurania'' , 2='' Melotrianthus'' , 3={{clade , 1=''

'' , 2={{clade , 1={{clade , 1={{clade , 1=''

'' , 2='' Halosicyos'' , 2=''

'' , 2={{clade , 1='' Tumamoca'' , 2=''

'' , targetB=CUCURBITEAE , subcladeB={{clade , 1={{clade , 1='' Polyclathra'' , 2={{clade , 1=''

'' , 2='' Cucurbita'' , 2={{clade , 1={{clade , 1='' Penelopeia'' , 2='' Calycophysum'' , 3='' Sicana'' , 2={{clade , 1='' Tecunumania'' , 2={{clade , 1='' Schizocarpum'' , 2={{clade , 1='' Cionosicys'' , 2={{clade , 1=''

'' , 2='' Cayaponia'' , targetC=BENINCASEAE , subcladeC={{clade , 1='' Zehneria'' , 2={{clade , 1={{clade , 1={{clade , 1='' Citrullus'' , 2={{clade , 1=''

'' , 2='' Lagenaria'' , 2={{clade , 1=''

'' , 2={{clade , 1={{clade , 1='' Raphidiocystis'' , 2={{clade , 1='' Cephalopentandra'' , 2={{clade , 1='' Lemurosicyos'' , 2={{clade , 1='' Solena'' , 2='' Borneosicyos'' , 2={{clade , 1='' Benincasa'' , 2={{clade , 1='' Ctenolepis'' , 2={{clade , 1=''

'' , 2='' Trochomeria'' , 2={{clade , 1={{clade , 1=''

'' , 2={{clade , 1='' Indomelothria'' , 2=''

'' , 2={{clade , 1={{clade , 1={{clade , 1=''

'' , 2='' Diplocyclos'' , 2={{clade , 1='' Papuasicyos'' , 2='' Scopellaria'' , 2={{clade , 1='' Muellerargia'' , 2='' Cucumis'' {{anchor, Pests, Diseases

Pests and diseases

Sweet potato whitefly is the vector of a number of cucurbit viruses that cause yellowing symptoms throughout the southern United States.{{cite journal , title=Whitefly-Transmitted Yellowing Viruses of Cucurbit Crops , publisher= American Phytopathological Society , date=2020-08-28 , doi=10.1094/grow-cuc-08-20-006 , first=William M. , last=Wintermantel, s2cid=242003450

References

{{Reflist

External links

Cucurbitaceae
i

i
L. Watson and M.J. Dallwitz (1992 onwards). The families of flowering plants: descriptions, illustrations, identification, information retrieval.
https://web.archive.org/web/20070103200438/http://delta-intkey.com/ {{Angiosperm families {{Taxonbar, from=Q8314 {{Authority control Rosid families Extant Danian first appearances Taxa named by Antoine Laurent de Jussieu Taxa described in 1789

Description

Fossil history

Classification

Tribal classification

Systematics

Pests and diseases

References

Further reading

External links