''Cryonectes'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

pliosaurid Pliosauridae is a family of plesiosaurian marine reptiles from the Latest Triassic to the early Late Cretaceous (Rhaetian to Turonian stages) of Australia, Europe, North America and South America. The family is more inclusive than the archetyp ...

plesiosauria The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...

n known from the

Early Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma ...

Normandy Normandy (; french: link=no, Normandie ; nrf, Normaundie, Nouormandie ; from Old French , plural of ''Normant'', originally from the word for "northman" in several Scandinavian languages) is a geographical and cultural region in Northwestern ...

, northern

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of overseas regions and territories in the Americas and the Atlantic, Pacific and Indian Oceans. Its metropolitan ar ...

Discovery

''Cryonectes'' is known only from the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

MAE Mae is an American rock band that formed in Norfolk, Virginia in 2001. The band's name is an acronym for "Multi-sensory Aesthetic Experience", based on a course taken by drummer Jacob Marshall while a student at Old Dominion University. History ...

2007.1.1(J), a partial

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...

and articulated

mandible In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bon ...

, and 10 associated

vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...

. It was collected in 2007 from the Calcaire à Bélemnites Formation of

, dating to the latest

Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 190.8 ± 1.5 Ma and 182.7 ± 1.5 Ma (million years ago). The ...

stage of the late

, about 185-183

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

. It was found in the Roche-Blain quarry, located in the commune of Fresney-le-Puceux near the city of Laize-la-Ville, south of

Caen Caen (, ; nrf, Kaem) is a commune in northwestern France. It is the prefecture of the department of Calvados. The city proper has 105,512 inhabitants (), while its functional urban area has 470,000,acid, which damaged some parts, especially the teeth, which are now broken and lack enamel. The complete

and the skull are in occlusion with teeth ''in situ'' and one separated and almost complete

tooth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, t ...

is also known. The

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has ...

e and

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...

e are partially preserved and the most important part of the palate is preserved separate from the skull. The ten associated vertebrae are preserved in a non-natural sequence, nine being

cervical In anatomy, cervical is an adjective that has two meanings: # of or pertaining to any neck. # of or pertaining to the female cervix: i.e., the ''neck'' of the uterus. *Commonly used medical phrases involving the neck are **cervical collar **cervic ...

, while the last one is indeterminable. Distinct neurocentral sutures are visible on all vertebrae preserving neural arches, except on the ninth vertebra, which presents a closed suture. Therefore, ''Cryonectes'' is at a relatively early stage in ontogenetic development. The ninth vertebra suggests that it was probably a subadult individual. ''Cryonectes'' is characterized by a unique combination of characters including a very slight constriction between premaxillae and maxillae, its snout is greatly elongated and the mandible has a long symphysis bearing seven tooth position and retaining a ventral mandibular ridge. A

cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived ch ...

analysis performed by Peggy Vincent, Nathalie Bardet and Emanuela Mattioli found it to be basal to ''

Pliosaurus ''Pliosaurus'' (meaning 'more lizard') is an extinct genus of thalassophonean pliosaurid known from the Kimmeridgian and Tithonian stages (Late Jurassic) of Europe and South America. Their diet would have included fish, cephalopods, and marine ...

'', ''

Peloneustes ''Peloneustes'' (meaning "mud swimmer") is a genus of pliosaurid plesiosaur from the Middle Jurassic of England. Its remains are known from the Peterborough Member of the Oxford Clay Formation, which is Callovian in age. It was originally de ...

'' and other pliosaurids.

Etymology

''Cryonectes'' was first named by Peggy Vincent, Nathalie Bardet and Emanuela Mattioli in

2012 File:2012 Events Collage V3.png, From left, clockwise: The passenger cruise ship Costa Concordia lies capsized after the Costa Concordia disaster; Damage to Casino Pier in Seaside Heights, New Jersey as a result of Hurricane Sandy; People gather ...

and the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

is ''Cryonectes neustriacus''. The generic name is derived from ''kryos'',

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

for "cold", and ''nektris'', for "swimmer", and refers to the cool climatic conditions that prevailed during the uppermost Pliensbachian. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

is derived from the Frank kingdom of Neustrie which covered northwest France, created after the death of Clovis I.

References

{{Taxonbar, from=Q5190542 Early Jurassic plesiosaurs of Europe Fossil taxa described in 2012 Fossils of France Sauropterygian genera

Discovery

Etymology

See also

References