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''Crotalus durissus'', known as the South American rattlesnake, tropical rattlesnake,Mehrtens JM. 1987. Living Snakes of the World in Color. New York: Sterling Publishers. 480 pp. and by other names, is a highly
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a ...
pit viper
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
found in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
. It is the most widely distributed member of its genus. Currently, seven subspecies are recognized.


Taxonomy

The Guiana rattlesnake, previously recognized as ''C. d. dryinus'', is now considered a synonym for ''C. d. durissus''. In fact, after the previous nominate subspecies for the ''C. d. durissus'' complex became the current nominate for ''
Crotalus simus : ''Common names: Middle American rattlesnake, Central American rattlesnake,''Klauber LM. 1997. Rattlesnakes: Their Habitats, Life Histories, and Influence on Mankind. Second Edition. 2 volumes. Reprint, University of California Press, Berkeley. ...
'', which now represents its Mexican and Central American members, ''C. d. dryinus'' became the new nominate for the South American rattlesnakes as represented by ''C. durissus''. The subspecies previously known as ''C. d. collilineatus'' and ''C. d. cascavella'' were moved to the synonymy of ''C. d. terrificus'' following the publication of a paper by Wüster ''et al.'' in 2005.


Subspecies


Description

A large
Neotropical The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In bi ...
rattlesnake, it grows to a length of , and rarely to a maximum length of . It has two distinct stripes starting at the base of the head. Within the lines, the color is lighter than the stripes. The color and pattern of the body are quite variable, most with an 18–32 dorsal with a darker diamond, and rhombic spots, 25–33 (usually 27) rows of dorsal scale in the middle of the body. The head has a dark brown bar at the top, with a dark post-orbital band. The color of the belly varies, it can be white or yellowish, with light gray spots, becoming darker towards the tail. The tail is usually gray, with dark and vague crossed bands.


Behaviour

The species is more active at dusk and in the early hours of the morning; it is usually not aggressive towards humans, but can bite if cornered or threatened.


Reproduction

The South American rattlesnake has a seasonal reproductive cycle; competition between males (for access to females) begins around the summer's end, with copulation occurring during the fall, and the birth of the young taking place the following spring/summer. Reproduction is
ovoviviparous Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
, giving birth to 4–8 young. In Roraima, Brazil it has been reported that the gestation lasts for 5 months, and they are capable of giving birth to up to 14 young.


Diet

The diet consists mainly of
rodents Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
, likely due to the great abundance and availability of these animals throughout the year, in most areas where the snakes reside. In some regions,
lizards Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia a ...
of the Teiidae family are also part of the diet of ''C. durissus''.


Common names

Common names for the species include: South American rattlesnake, tropical rattler, tropical rattlesnake, neotropical rattlesnake,U.S. Navy. 1991. Poisonous Snakes of the World. US Govt. New York: Dover Publications Inc. 203 pp. Guiana rattlesnake (previously used for ''C. d. dryinus'').Brown JH. 1973. Toxicology and Pharmacology of Venoms from Poisonous Snakes. Springfield, Illinois: Charles C. Thomas. 184 pp. LCCCN 73-229. and in
Spanish Spanish might refer to: * Items from or related to Spain: ** Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Ca ...
: ''víbora de cascabel, cascabel, cascabela'', and also in Portuguese, ''cascavel''. In Suriname it is known as Sakasneki.


Geographic range

''Crotalus durissus'' is found in South America except the Andes Mountains. However, its range is discontinuous, with many isolated populations in northern South America, including Colombia,
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in ...
, Guyana,
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the north ...
,
French Guiana French Guiana ( or ; french: link=no, Guyane ; gcr, label=French Guianese Creole, Lagwiyann ) is an overseas departments and regions of France, overseas department/region and single territorial collectivity of France on the northern Atlantic ...
and northern
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. It occurs in Colombia and eastern Brazil to southeastern
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = Seal (emblem), National seal , national_motto = "Fi ...
,
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
,
Paraguay Paraguay (; ), officially the Republic of Paraguay ( es, República del Paraguay, links=no; gn, Tavakuairetã Paraguái, links=si), is a landlocked country in South America. It is bordered by Argentina to the south and southwest, Brazil to th ...
,
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
, and northern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
( Catamarca,
Córdoba Córdoba most commonly refers to: * Córdoba, Spain, a major city in southern Spain and formerly the imperial capital of Islamic Spain * Córdoba, Argentina, 2nd largest city in the country and capital of Córdoba Province Córdoba or Cordoba may ...
,
Corrientes Corrientes (; Guaraní: Taragüí, literally: "Currents") is the capital city of the province of Corrientes, Argentina, located on the eastern shore of the Paraná River, about from Buenos Aires and from Posadas, on National Route 12. It has ...
, Chaco, Entre Rios,
Formosa Taiwan, officially the Republic of China (ROC), is an island country located in East Asia. The main island of Taiwan, formerly known in the Western political circles, press and literature as Formosa, makes up 99% of the land area of the territori ...
, La Pampa,
La Rioja La Rioja () is an autonomous community and province in Spain, in the north of the Iberian Peninsula. Its capital is Logroño. Other cities and towns in the province include Calahorra, Arnedo, Alfaro, Haro, Santo Domingo de la Calzada, a ...
, Mendoza,
Misiones Misiones (, ''Missions'') is one of the 23 provinces of Argentina, located in the northeastern corner of the country in the Mesopotamia region. It is surrounded by Paraguay to the northwest, Brazil to the north, east and south, and Corrientes P ...
,
San Juan San Juan, Spanish for Saint John, may refer to: Places Argentina * San Juan Province, Argentina * San Juan, Argentina, the capital of that province * San Juan, Salta, a village in Iruya, Salta Province * San Juan (Buenos Aires Underground), ...
,
San Luis San Luis (Spanish for "Saint Louis") may refer to: Places Argentina * San Luis Province * San Luis, Argentina, capital of San Luis Province Belize * San Luis, Belize, in Orange Walk District Colombia * San Luis, Antioquia, a town and municipality ...
, Santa Fe,
Santiago del Estero Santiago del Estero (, Spanish for ''Saint-James-Upon-The-Lagoon'') is the capital of Santiago del Estero Province in northern Argentina. It has a population of 252,192 inhabitants, () making it the twelfth largest city in the country, with a su ...
and Tucumán). It also occurs on some islands in the Caribbean, including Aruba. The
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
given is "America."


Habitat

It prefers
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
and
semi-arid A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi- ...
zones. It has been reported to occur in
littoral The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal areas ...
xerophilous
scrub Scrub(s) may refer to: * Scrub, low shrub and grass characteristic of scrubland * Scrubs (clothing), worn by medical staff * ''Scrubs'' (TV series), an American television program * Scrubs (occupation), also called "scrub tech," "scrub nurse," ...
, psammophilous and halophilous littoral
grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
, thorny xerophilous scrub, tropophilous
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, a ...
and semideciduous scrub, as well as tropophilous semideciduous seasonal forest in the northwest of
Venezuela Venezue