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''Crotalus cerberus'' is a
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
pit viper The Crotalinae, commonly known as pit vipers,Mehrtens JM (1987). ''Living Snakes of the World in Color''. New York: Sterling Publishers. 480 pp. . or pit adders, are a subfamily (biology), subfamily of Viperidae, vipers found in Asia and the ...
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
found in the southwestern
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
. It is known as the Arizona black rattlesnake, black rattlesnake, and several other common names. Wright AH, Wright AA. 1957. Handbook of Snakes. 2 volumes. Comstock Publishing Associates. (7th printing, 1985). 1105 pp. .


Description

According to Wright and Wright (1957), adults grow to an average length of . Klauber (1997) reports the maximum length to be less at , with the smallest gravid female measuring . Klauber LM. 1997. Rattlesnakes: Their Habitats, Life Histories, and Influence on Mankind. Second Edition. 2 volumes. Reprint, University of California Press, Berkeley. . Hubbs and O'Connor (2012) list an adult size range of .Hubbs, Brian and Brendan O'Connor. 2012. '' A Guide to the Rattlesnakes and other Venomous Serpents of the United States''. Tricolor Books. Tempe, Arizona, 129 pp. . (''Crotalus cerberus'', pp. 34-35.) The color pattern consists of a dark grayish, brownish black, reddish brown, or blackish ground color, overlaid with a dorsal pattern of blotches that are rectangular anteriorly, becoming subhexagonal posteriorly, eventually becoming crossbands just before the tail. However, specimens also may be a uniform dark color without any clear dorsal pattern, or the dorsal blotches may be even darker and bordered with white, cream, or yellow transverse rows of scales, or the color pattern may be quite pale with a significant amount of yellow mixed in. A postocular stripe is evident in lightly colored specimens, but not so much in darker ones.Campbell JA, Lamar WW. 2004. The Venomous Reptiles of the Western Hemisphere. 2 volumes. Comstock Publishing Associates, Ithaca and London. 870 pp. 1500 plates. .


Color change

Arizona black rattlesnakes undergo morphological color change (gradual change due to changes in number or quality of chromatophores); adults are less patterned than juveniles (also called ontogenetic color change). Some adults can change color relatively quickly (physiological color change: rapid change due to movement of organelles within chromatophores), an ability shared not only with
chameleon Chameleons or chamaeleons (Family (biology), family Chamaeleonidae) are a distinctive and highly specialized clade of Old World lizards with 200 species described as of June 2015. The members of this Family (biology), family are best known for ...
s and other lizards but also with other snakes such as some
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Tropidophis ''Tropidophis'', common name Caribbean dwarf boas, wood snakes or West Indian wood snakes,Hampton Wildman Parker, Parker HW, species:Alice Georgie Cruickshank Grandison, Grandison AGC (1977). ''Snakes — a Natural History''. Second Edition. Lond ...
''. The mechanism for their physiological color change is likely the same as that documented in the closely related prairie rattlesnake (melanin movement within their dermal melanophores), but further research is needed to determine the stimuli for this phenomenon in this rattlesnake.


Common names

Arizona black rattlesnake, black rattlesnake, black diamond rattlesnake, brown rattlesnake, Cerberus rattlesnake, mountain diamond-back. Also often incorrectly referred to as a timber rattlesnake.


Geographic range

Found in the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, in
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
from the
Hualapai Mountains The Hualapai Mountains are a mountain range located in Mohave County, Arizona, Mohave County, east of Kingman, Arizona, Kingman, Arizona. Rising up to 8,417 feet at its highest peak, the higher elevations of the Hualapai Mountains support Madrea ...
and Cottonwood Cliffs in the northwest of the state, southeast to the Santa Catalina, Rincon, Pinaleno and Blue Mountains. Also found at Steeple Rock, in extreme western
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
. The type locality given is "San Francisco Mountains" (
Coconino County, Arizona Coconino County is a County (United States), county in the North Central Arizona, North-Central part of the U.S. state of Arizona. Its population was 145,101 at the 2020 United States census, 2020 census. The county seat is Flagstaff, Arizon ...
, USA).


Diet

It preys upon suitably sized
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s,
reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
s,
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s and their
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
s, and
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s.


Reproduction

Sexually mature females bear live young in broods of 4 to 21
neonate In common terminology, a baby is the very young offspring of adult human beings, while infant (from the Latin word ''infans'', meaning 'baby' or 'child') is a formal or specialised synonym. The terms may also be used to refer to Juvenile (orga ...
s. The Arizona black rattlesnake is the first species of snake observed to exhibit complex social behavior, and like all temperate pit vipers, care for their babies. Females remain with their young in nests for 7 to 14 days, and mothers have been observed cooperatively parenting their broods.


References


External links


Arizona Black Rattlesnake, ''Crotalus cerberus''
at th

Accessed 18 June 2008. {{Taxonbar, from1=Q3003829, from2=Q5188959
cerberus In Greek mythology, Cerberus ( or ; ''Kérberos'' ), often referred to as the hound of Hades, is a polycephaly, multi-headed dog that guards the gates of the Greek underworld, underworld to prevent the dead from leaving. He was the offspring o ...
Snakes of North America Endemic reptiles of the United States Fauna of the Southwestern United States Fauna of the Sonoran Desert Natural history of Arizona Natural history of New Mexico Taxa named by Elliott Coues Reptiles described in 1875