A crossbreed is an organism with
purebred
Purebreds are cultivars of an animal species achieved through the process of selective breeding. When the lineage of a purebred animal is recorded, that animal is said to be pedigreed. Purebreds breed true-to-type, which means the progeny of l ...
parents of two different breeds, varieties, or populations. A domestic animal of unknown ancestry, where the breed status of only one parent or grandparent is known, may also be called a crossbreed though the term "mixed breed" is technically more accurate.
Outcrossing
Out-crossing or out-breeding is the technique of crossing between different breeds. This is the practice of introducing distantly related genetic material into a breeding line, thereby increasing genetic diversity.
Outcrossing in animals
Out ...
is a type of crossbreeding used within a purebred breed to increase the genetic diversity within the breed, particularly when there is a need to avoid
inbreeding
Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely genetic distance, related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genet ...
.
In animal breeding, ''crossbreeds'' are crosses within a single species, while ''
hybrids'' are crosses between different species. In plant breeding terminology, the term ''crossbreed'' is uncommon, and no universal term is used to distinguish hybridization or crossing within a population from those between populations, or even those between species.
Crossbreeding is the process of breeding such an organism. It can be beneficially used to maintain health and viability of organisms. However, irresponsible crossbreeding can also produce organisms of inferior quality or dilute a
purebred
Purebreds are cultivars of an animal species achieved through the process of selective breeding. When the lineage of a purebred animal is recorded, that animal is said to be pedigreed. Purebreds breed true-to-type, which means the progeny of l ...
gene pool to the point of
extinction
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
of a given
breed
A breed is a specific group of breedable domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist seve ...
of organism.
Crossbreeds in specific animals
Cats: The many newly developed and recognized
breeds of domestic cat are crossbreeds between existing, well-established breeds (sometimes with limited
hybridization with some wild species), to either combine selected traits from the
foundation stock
Foundation stock or foundation bloodstock refers to animals that are the progenitors, or foundation, of a breed or of a given bloodline within such. Many modern breeds can be traced to specific, named foundation animals, but a group of animals may ...
, or propagate a rare
mutation
In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA or viral replication, ...
without excessive
inbreeding
Inbreeding is the production of offspring from the mating or breeding of individuals or organisms that are closely genetic distance, related genetically. By analogy, the term is used in human reproduction, but more commonly refers to the genet ...
. However, some nascent breeds such as the
Aegean cat
Aegean cats ( ''gáta tou Aigaíou'') are a naturally occurring landrace of domestic cat originating from the Cycladic Islands of Greece and western Turkey. It is considered a natural cat, developing without human interference. Development of th ...
are developed entirely from a local
landrace
A landrace is a Domestication, domesticated, locally adapted, often traditional variety of a species of animal or plant that has developed over time, through adaptation to its natural and cultural Environment (biophysical), environment of agric ...
population. Most
experimental cat breeds are crossbreeds.
Cattle: In cattle, there are systems of crossbreeding. In many crossbreeds, one animal is larger than the other. One is used when the purebred females are particularly adapted to a specific environment, and are crossed with purebred bulls from another environment to produce a generation having traits of both parents.
Sheep: The large number of breeds of sheep, which vary greatly, creates an opportunity for crossbreeding to be used to tailor production of lambs to the goal of the individual stockman.
Llamas: Results of crossbreeding classic and woolly breeds of llama are unpredictable. The resulting offspring displays physical characteristics of either parent, or a mix of characteristics from both, periodically producing a fleeced llama.
The results are increasingly unpredictable when both parents are crossbreeds, with possibility of the offspring displaying characteristics of a grandparent, not obvious in either parent.
Dogs:
A ''crossbred'' dog is a cross between two (sometimes more) known breeds, and is usually distinguished from a ''
mixed-breed dog
A mongrel, mutt, or mixed-breed dog is a dog that does not belong to one officially recognized Dog breed, breed, including those that result from intentional Dog breeding, breeding. Although the term ''mixed-breed dog'' is sometimes preferred, m ...
'', which has ancestry from many sources, some of which may not be known. Crossbreeds are popular, due to the belief that they have
increased vigor without loss of attractiveness of the dog. Certain planned crossbreeding between purebred dogs of different breeds are now widely known as "designer dogs" and can produce puppies worth more than their purebred parents, due to a high demand.

Horses:
Crossbreeding horses is often done with the intent of ultimately creating a new breed of horse. One type of modern crossbreeding in horses created many of the
warmblood
Warmbloods are a group of middle-weight horse types and breeds primarily originating in Europe and registered with organizations that are characterized by open studbook policy, studbook selection, and the aim of breeding for equestrian sp ...
breeds used in the
sport horse disciplines, usually registered in an
open stud book
A breed registry, also known as a herdbook, studbook or register, in animal husbandry, the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeders w ...
by a
studbook selection
Studbook selection is a process used in certain breeds of horses to select breeding stock. It allows a breed registry to direct the evolution of the breed towards the ideal by eliminating unhealthy or undesirable animals from the population. The r ...
procedure that evaluates conformation, pedigree and, in some animals, a training or performance standard. Most warmblood breeds began as a cross of
draft horse
A draft horse (US) or draught horse (UK), also known as dray horse, carthorse, work horse or heavy horse, is a large horse bred to be a working animal hauling freight and doing heavy agricultural tasks such as plowing. There are a number o ...
breeds on
Thoroughbred
The Thoroughbred is a list of horse breeds, horse breed developed for Thoroughbred racing, horse racing. Although the word ''thoroughbred'' is sometimes used to refer to any breed of purebred horse, it technically refers only to the Thorough ...
s, but have, in some cases, developed over the past century to the point where they are considered to be a true-breeding population and have a
closed stud book
Closed may refer to:
Mathematics
* Closure (mathematics), a set, along with operations, for which applying those operations on members always results in a member of the set
* Closed set, a set which contains all its limit points
* Closed interval, ...
. Other types of recognized crossbreeding include that within the
American Quarter Horse
The American Quarter Horse, or Quarter Horse, is an American breed of horse that excels at sprinting short distances. Its name is derived from its ability to outrun other horse breeds in races of or less; some have been clocked at speeds up to ...
, which will register horses with one Thoroughbred parent and one registered Quarter Horse parent in the "Appendix" registry, and allow such animals full breed registration status as Quarter Horses if they meet a certain performance standard. Another well-known crossbred horse is the
Anglo-Arabian, which may be produced by a purebred
Arabian horse
The Arabian or Arab horse ( , DIN 31635, DMG ''al-ḥiṣān al-ʿarabī'') is a horse breed, breed of horse with historic roots on the Arabian Peninsula. With a distinctive head shape and high tail carriage, the Arabian is one of the most easi ...
crossed on a Thoroughbred, or by various crosses of Anglo-Arabians with other Anglo-Arabians, as long as the ensuing animal never has more than 75% or less than 25% of each breed represented in its pedigree.
Hybrid animals
A
hybrid animal is one with parentage of two separate species, differentiating it from crossbred animals, which have parentage of the same species. Hybrids are usually, but not always, sterile.
One of the most ancient types of hybrid animal is the
mule
The mule is a domestic equine hybrid between a donkey, and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two ...
, a cross between a female
horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
and a male
donkey
The donkey or ass is a domesticated equine. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a separate species, ''Equus asinus''. It was domes ...
. The
liger
The liger is a hybrid (biology), hybrid offspring of a male lion (''Panthera leo'') and a tigress, or female tiger (''Panthera tigris''). The liger has parents in the same genus but of different species. The liger is distinct from the opposite ...
is a
hybrid cross between a male
lion
The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...
and female
tiger
The tiger (''Panthera tigris'') is a large Felidae, cat and a member of the genus ''Panthera'' native to Asia. It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly vertical stripes. It is ...
. The
yattle is a cross between a
cow and a
yak. Other crosses include the
tigon
The tigon is a hybrid offspring of a male tiger (''Panthera tigris'') and a female lion, or lioness (''Panthera leo'').[yakalo
The yakalo is a cross of the yak (''Bos grunniens'') and the American bison (''Bison bison'', known as a buffalo in North America). It was produced by hybridisation experiments in the 1920s, when crosses were made between yak bulls and both pur ...](_blank)
(between a yak and an
American bison
The American bison (''Bison bison''; : ''bison''), commonly known as the American buffalo, or simply buffalo (not to be confused with Bubalina, true buffalo), is a species of bison that is endemic species, endemic (or native) to North America. ...
). The Incas recognized that hybrids of ''Lama glama'' (llama) and ''Vicugna pacos'' (alpaca) resulted in a hybrid with none of the advantages of either parent.
At one time it was thought that dogs and wolves were separate species, and the
crosses between dogs and wolves were called wolf hybrids. Today wolves and dogs are both recognized as ''
Canis lupus
The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, including the dog and dingo, though gr ...
'', but the old term "wolf hybrid" is still used.
Mixed breeds
A mixed-breed animal is defined as having undocumented or unknown parentage, while a crossbreed generally has known, usually
purebred
Purebreds are cultivars of an animal species achieved through the process of selective breeding. When the lineage of a purebred animal is recorded, that animal is said to be pedigreed. Purebreds breed true-to-type, which means the progeny of l ...
parents of two distinct breeds or varieties. A dog of unknown parentage is often called a mixed-breed dog, "mutt" or "
mongrel
A mongrel, mutt, or mixed-breed dog is a dog that does not belong to one officially recognized breed, including those that result from intentional breeding. Although the term ''mixed-breed dog'' is sometimes preferred, many mongrels have no kn ...
." A cat of unknown parentage is often referred to as a
domestic short-haired or
domestic long-haired cat generically, and in some dialects is often called a "moggie". A horse of unknown bloodlines is called a
grade horse.
Designer crossbreed
A designer crossbreed or designer breed is a crossbred animal with
purebred
Purebreds are cultivars of an animal species achieved through the process of selective breeding. When the lineage of a purebred animal is recorded, that animal is said to be pedigreed. Purebreds breed true-to-type, which means the progeny of l ...
parents, usually registered with a
breed registry
A breed registry, also known as a herdbook, studbook or register, in animal husbandry, the hobby of animal fancy, is an official list of animals within a specific breed whose parents are known. Animals are usually registered by their breeders w ...
, but from two different
breed
A breed is a specific group of breedable domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist seve ...
s. These animals are the result of a deliberate decision to create a specific crossbred animal.
Less often, the animal may have more than two pure breeds in its ancestry, but unlike a mutt or a
mongrel
A mongrel, mutt, or mixed-breed dog is a dog that does not belong to one officially recognized breed, including those that result from intentional breeding. Although the term ''mixed-breed dog'' is sometimes preferred, many mongrels have no kn ...
, its entire
pedigree is known to descend from specific known animals. While the term is best known when applied to certain
dog crossbreed
Dog crossbreeds (sometimes called designer dogs) are dogs which have been Artificial selection, intentionally bred from two or more recognized dog breeds. They are not Mongrel#Crossbreed vs. mongrel, dogs with no purebred ancestors, but are not ...
s, other animals such as cattle, horses, birds and cats may also be bred in this fashion. Some crossbred breeders start a freestanding breed registry to record designer crossbreds, other crossbreds may be included in an "appendix" to an existing purebred registry. either form of registration may be the first step in recording and tracking
pedigrees in order to develop a new breed.
The purpose of creating designer crossbreds is usually one or more of the following reasons:
# to breed animals with
heterosis
Heterosis, hybrid vigor, or outbreeding enhancement is the improved or increased function of any biological quality in a hybrid offspring. An offspring is heterotic if its traits are enhanced as a result of mixing the genetic contributions o ...
, commonly known as "hybrid vigor",
# to create animals with more predictable characteristics than
mixed breed
A mixed breed is a domesticated animal descended from multiple breeds of the same species, often breeding without any human intervention, recordkeeping, or selective breeding. Examples include:
* Mixed-breed dog
A mongrel, mutt, or mixed ...
or
mongrel
A mongrel, mutt, or mixed-breed dog is a dog that does not belong to one officially recognized breed, including those that result from intentional breeding. Although the term ''mixed-breed dog'' is sometimes preferred, many mongrels have no kn ...
breeding,
# to avoid certain undesirable
recessive
In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and ...
traits that lead to
genetic disease
A genetic disorder is a health problem caused by one or more abnormalities in the genome. It can be caused by a mutation in a single gene (monogenic) or multiple genes (polygenic) or by a chromosome abnormality. Although polygenic disorders are ...
s that plague many purebred animals,
# to develop an animal that combines what are viewed as the best traits of two or more breeds,
# as the preliminary steps toward developing a new animal breed.
Breeders of designer crossbreds borrow the
technical language from
hybrid plant breeding: A first generation, 50–50 crossbred is an
F1 cross.
Subsequent generations may see a purebred animal crossed back on a crossbred, creating a 75/25 cross,
or a BC1 or F1b "
backcross." The breeding of two crossbreeds of the same combination of breeds, creating an F2 cross, an animal that is still a 50–50 cross, but it is the second
filial generation of the combination.
An F2 cross bred to an F2 cross creates an F3 cross. Similarly, an F2 animal bred to an F1 animal creates an F2b backcross. F3 crosses and greater are called "multi-generational" crosses. In dog breeding, three generations of reliable documented breeding can be considered a "breed" rather than a crossbreed.
There are disadvantages to creating designer crossbreeds, notably the potential that the cross will be of inferior quality or that it will not produce as consistent a result as would breeding purebred animals. For example, the
Poodle
The Poodle, called the in German () and the in French, is a breed of water dog. The breed is divided into four varieties based on size, the Standard Poodle, Medium Poodle, Miniature Poodle and Toy Poodle, although the Medium Poodle is no ...
is a frequent breed used in creation of designer crossbreeds, due to its non-shedding coat, but that trait does not always breed true when it is part of a designer cross.
Also, because breeders of crossbred animals may be less careful about
genetic testing
Genetic testing, also known as DNA testing, is used to identify changes in DNA sequence or chromosome structure. Genetic testing can also include measuring the results of genetic changes, such as RNA analysis as an output of gene expression, or ...
and weeding out undesirable traits,
certain deleterious
dominant genes may still be passed on to a crossbreed offspring. In an F2 cross, recessive genetic traits may also return if the parent animals were both carriers of an undesired trait.
See also
*
Artificial selection
Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant ...
*
Canid hybrid
Canid hybrids are the result of interbreeding between the species of the subfamily ''Caninae''.
Genetic considerations
The wolf-like canids are a group of large carnivores that are genetically closely related because they all possess 78 chromos ...
*
Heterosis
Heterosis, hybrid vigor, or outbreeding enhancement is the improved or increased function of any biological quality in a hybrid offspring. An offspring is heterotic if its traits are enhanced as a result of mixing the genetic contributions o ...
*
Introgression
Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Introg ...
*
Selective breeding
Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant m ...
References
{{Authority control
Animal breeding
Breeding