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The crop (also the croup, the craw, the ingluvies, and the sublingual pouch) is a thin-walled, expanded portion of the
alimentary tract The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The tract is the largest of the body's systems, after the cardiovascular system. T ...
, which is used for the storage of food before
digestion Digestion is the breakdown of large insoluble food compounds into small water-soluble components so that they can be absorbed into the blood plasma. In certain organisms, these smaller substances are absorbed through the small intestine into th ...
. The crop is an anatomical structure in vertebrate animals, such as
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s, and invertebrate animals, such as
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
s (snails and slugs),
earthworm An earthworm is a soil-dwelling terrestrial invertebrate that belongs to the phylum Annelida. The term is the common name for the largest members of the class (or subclass, depending on the author) Oligochaeta. In classical systems, they we ...
s,
leech Leeches are segmented parasitism, parasitic or Predation, predatory worms that comprise the Class (biology), subclass Hirudinea within the phylum Annelida. They are closely related to the Oligochaeta, oligochaetes, which include the earthwor ...
es, and
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s.


Insects

Cropping is used by
bees Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...
to temporarily store nectar of flowers. When bees "suck" nectar, it is stored in their crops. Other
Hymenoptera Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typi ...
also use crops to store liquid food. The crop in eusocial insects, such as
ants Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of ...
, has specialized to be distensible, and this specialization enables important communication between colonial insects through
trophallaxis Trophallaxis () is the transfer of food or other fluids among members of a community through mouth-to-mouth (stomodeum, stomodeal) or anus-to-mouth (proctodeum, proctodeal) feeding. Along with nutrients, trophallaxis can involve the transfer of m ...
. The crop can be found in the
foregut The foregut in humans is the anterior part of the alimentary canal, from the distal esophagus to the first half of the duodenum, at the entrance of the bile duct. Beyond the stomach, the foregut is attached to the abdominal walls by mesentery. ...
of insects.


Birds

In a
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
's digestive system, the crop is an expanded, muscular pouch near the gullet or throat. It is a part of the digestive tract, essentially an enlarged part of the
esophagus The esophagus (American English), oesophagus (British English), or œsophagus (Œ, archaic spelling) (American and British English spelling differences#ae and oe, see spelling difference) all ; : ((o)e)(œ)sophagi or ((o)e)(œ)sophaguses), c ...
. As with most other organisms that have a crop, it is used to temporarily store food. Not all bird species have one. In adult doves and pigeons, it can produce crop milk to feed newly hatched birds. Scavenging birds, such as
vulture A vulture is a bird of prey that scavenges on carrion. There are 23 extant species of vulture (including condors). Old World vultures include 16 living species native to Europe, Africa, and Asia; New World vultures are restricted to Nort ...
s, will gorge themselves when prey is abundant, causing their crop to bulge. They subsequently sit, sleepy or half torpid, to digest their food. Most raptors, including
hawk Hawks are birds of prey of the family Accipitridae. They are very widely distributed and are found on all continents, except Antarctica. The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks, and others. This ...
s,
eagle Eagle is the common name for the golden eagle, bald eagle, and other birds of prey in the family of the Accipitridae. Eagles belong to several groups of Genus, genera, some of which are closely related. True eagles comprise the genus ''Aquila ( ...
s and
vulture A vulture is a bird of prey that scavenges on carrion. There are 23 extant species of vulture (including condors). Old World vultures include 16 living species native to Europe, Africa, and Asia; New World vultures are restricted to Nort ...
s (as stated above), have a crop; however,
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s do not. Similarly, all true quail ( Old World quail and
New World quail The New World quail are small birds, that despite their similar appearance and habits to the Old World quail, belong to a different family known as the Odontophoridae. In contrast, the Old World quail are in the Phasianidae family, sharing only a ...
) have a crop, but buttonquail do not.
Chicken The chicken (''Gallus gallus domesticus'') is a domesticated subspecies of the red junglefowl (''Gallus gallus''), originally native to Southeast Asia. It was first domesticated around 8,000 years ago and is now one of the most common and w ...
s,
turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
s, ducks and geese possess a crop, as do
parrot Parrots (Psittaciformes), also known as psittacines (), are birds with a strong curved beak, upright stance, and clawed feet. They are classified in four families that contain roughly 410 species in 101 genus (biology), genera, found mostly in ...
s. Pigeons also have crops; one domestic breed type is even bred to exaggerate the typical crop-inflating behavior so that the crop is inflated like a
balloon A balloon is a flexible membrane bag that can be inflated with a gas, such as helium, hydrogen, nitrous oxide, oxygen, or air. For special purposes, balloons can be filled with smoke, liquid water, granular media (e.g. sand, flour or rice), ...
. Some extinct birds like
Enantiornithes The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct Avialae, avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teet ...
did not have crops.


Literary references

In the
Sherlock Holmes Sherlock Holmes () is a Detective fiction, fictional detective created by British author Arthur Conan Doyle. Referring to himself as a "Private investigator, consulting detective" in his stories, Holmes is known for his proficiency with obser ...
story " The Adventure of the Blue Carbuncle" (1892), a valuable gem is hidden inside a bird's crop. "Craw" is an obsolete term for "crop", and this is still seen in the saying "it sticks in my craw" meaning "I can't etaphoricallyswallow it", that is, that a situation or other entity is unacceptable, or at any rate annoying.


See also

*
Esophagus The esophagus (American English), oesophagus (British English), or œsophagus (Œ, archaic spelling) (American and British English spelling differences#ae and oe, see spelling difference) all ; : ((o)e)(œ)sophagi or ((o)e)(œ)sophaguses), c ...
*
Gizzard The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including archosaurs (birds and other dinosaurs, crocodiles, alligators, pterosaurs), earthworms, some gast ...
* Gular pouch, in
bird anatomy Bird anatomy, or the physiological structure of birds' bodies, shows many unique adaptations, mostly aiding flight. Birds have a light skeletal system and light but powerful musculature which, along with circulatory and respiratory systems capa ...
, a flap generally used to store fish and other prey while hunting *
Intestines The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The tract is the largest of the body's systems, after the cardiovascular system. ...
*
Proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects. Birds Th ...
*
Stomach The stomach is a muscular, hollow organ in the upper gastrointestinal tract of Human, humans and many other animals, including several invertebrates. The Ancient Greek name for the stomach is ''gaster'' which is used as ''gastric'' in medical t ...


References


External links


The Alimentary Canal in Birds
{{Gastropod anatomy Annelid anatomy Bird anatomy Gastropod anatomy Insect anatomy Dinosaur anatomy