''Crocodylus anthropophagus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include ...

from the

Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...

Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...

. It lived 1.84 million years ago. It was a large-sized

predator Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common List of feeding behaviours, feeding behaviours that includes parasitism and micropredation ...

reaching a length of .

Etymology

''Crocodylus anthropophagus'' was first named by Christopher A. Brochu, Jackson Njau, Robert J. Blumenschine and Llewellyn D. Densmore in

2010 The year saw a multitude of natural and environmental disasters such as the 2010 Haiti earthquake, the Deepwater Horizon oil spill, and the 2010 Chile earthquake. The 2009 swine flu pandemic, swine flu pandemic which began the previous year ...

. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''anthropophagus'' is from Greek word "''anthropos''" that means "human" and Greek word "'" that means "eater", in reference to the evidence that this animal included hominids in its diet.

Taxonomy

The

holotype specimen A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was Species description, formally described. It is either the single such physical example (or illus ...

, NNHM-1001, comprises a skull and partial skeleton. All specimens were discovered in

Olduvai Gorge The Olduvai Gorge or Oldupai Gorge in Tanzania is one of the most important paleoanthropology, paleoanthropological localities in the world; the many sites exposed by the gorge have proven invaluable in furthering understanding of early human evo ...

, Tanzania, in two different rock formations dating to 1.845 and 1.839 million years ago in the

Plio-Pleistocene The Plio-Pleistocene is an informally described geological pseudo-period, which begins about 5 million years ago (Mya) and, drawing forward, combines the time ranges of the formally defined Pliocene and Pleistocene epochs—marking from about 5&n ...

. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below is from a 2021 study based on the finding of a new ''C. anthropophagus'' partial

cranium The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

Description

The skeleton is consistent with living representatives of the genus. The

axis An axis (: axes) may refer to: Mathematics *A specific line (often a directed line) that plays an important role in some contexts. In particular: ** Coordinate axis of a coordinate system *** ''x''-axis, ''y''-axis, ''z''-axis, common names ...

vertebra has a projection on the underside (a hypapophysis) which seems to have largely fused to the vertebra itself. The vertebrae are procoelous in shape, with a spherical projection extending from the back side which attaches to a concave socket on the front end of the preceding vertebra. The femora slightly bend in an S-shape. However, ''C. anthropophagus'' lacks the shallow bony pair of crests running from the eyes to the nose like in Indopacific ''Crocodylus'', and lacks a boss (a lump of bone) on the middle of the snout like in Neotropical ''Crocodylus''. The right

premaxilla The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammals h ...

(the bone that makes up the front end of the snout) of the holotype preserved three tooth sockets, with a notable gap in between the first and second sockets. Another specimen's left

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

(the bone that makes up the back end of the snout) preserved 13 circular sockets, though it may have partially preserved a 14th. There was a gap between the 9th and 10th. The teeth were all conical and lacked serrations. It had two prominent, triangular “horns” over the ears.

Paleoecology

''Crocodylus anthropophagus'' was the largest predator encountered by human ancestors at

, as indicated by

hominin The Hominini (hominins) form a taxonomic tribe of the subfamily Homininae (hominines). They comprise two extant genera: ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos), and in standard usage exclude the genus '' Gorilla'' ( gorillas) ...

specimens preserving crocodile bite marks from these sites. Its type locality is near those for ''

Homo habilis ''Homo habilis'' ( 'handy man') is an extinct species of archaic human from the Early Pleistocene of East and South Africa about 2.4 million years ago to 1.65 million years ago ( mya). Upon species description in 1964, ''H. habilis'' was highly ...

'' and ''

Paranthropus boisei ''Paranthropus boisei'' is a species of australopithecine from the Early Pleistocene of East Africa about 2.5 to 1.15 million years ago. The holotype specimen, OH 5, was discovered by palaeoanthropologist Mary Leakey in 1959 at Olduvai Gorge, Ta ...

''.

References

This article incorporates CC-BY-2.5 text from the reference.

External links

Ancient Human Ancestors Faced Fearsome Horned Crocodile
LiveScience {{Taxonbar, from=Q1932167 Crocodylidae Reptiles described in 2010 Fossil taxa described in 2010 Pleistocene crocodylomorphs Pliocene crocodylomorphs Pleistocene reptiles of Africa Pliocene reptiles