Crax Rubra
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The great curassow (''Crax rubra'') is a large,
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera's native range is restricted to Eura ...
-like bird from the
Neotropic The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeogra ...
al
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s, its range extending from eastern Mexico, through Central America to western Colombia and northwestern Ecuador. Male birds are black with curly crests and yellow beaks; females come in three colour morphs, barred, rufous and black. These birds form small groups, foraging mainly on the ground for fruits and
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s, and the occasional small
vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...
, but they roost and nest in trees. This species is
monogamous Monogamy ( ) is a relationship of two individuals in which they form a mutual and exclusive intimate partnership. Having only one partner at any one time, whether for life or serial monogamy, contrasts with various forms of non-monogamy (e.g. ...
, the male usually building the rather small nest of leaves in which two eggs are laid. This species is threatened by loss of habitat and hunting, and the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
has rated its conservation status as " vulnerable".


Taxonomy

The great curassow is the most northerly ''Crax'' species. It is part of a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
that inhabited the north of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
since about 9 mya (
Tortonian The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Tort ...
, Late
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
). As the
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
n
Andes The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the List of longest mountain chains on Earth, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range ...
were uplifted around 6 mya, this species' ancestors were cut off from the population to their southeast. The latter would in time evolve into the blue-billed curassow. The ancestral great curassows then spread along the Pacific side of the Andes, and into Central America during the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
as part of the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which land ...
.


Description

At in length and in weight, this is a very large cracid. Females are somewhat smaller than males. It is the most massive and heavy species in the family but its length is matched by a few other cracids. Three other species of curassow (the northern helmeted, the southern helmeted, and the
black Black is a color that results from the absence or complete absorption of visible light. It is an achromatic color, without chroma, like white and grey. It is often used symbolically or figuratively to represent darkness.Eva Heller, ''P ...
are all around the same average length as the great curassow. In this species, standard measurements are as follows: the wing chord is , the
tail The tail is the elongated section at the rear end of a bilaterian animal's body; in general, the term refers to a distinct, flexible appendage extending backwards from the midline of the torso. In vertebrate animals that evolution, evolved to los ...
is and the tarsus is . They have the largest mean standard measurements in the family, other than tail length. The male is black with a curly crest, a white belly, and a yellow knob on its bill. There are three morphs of female great curassows: barred morph females with barred neck, mantle, wings and tail; rufous morph with an overall reddish brown
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
and a barred tail; and dark morph female with a blackish neck, mantle and tail (the tail often faintly vermiculated), and some barring to the wings. In most regions only one or two morphs occur, and females showing a level of intermediacy between these morphs are known (e.g. resembling rufous morph, but with black neck and faint vermiculations to the wings). This species has a similar voice to several other curassows, its call consisting of a "peculiar" lingering whistle.


Behavior and ecology


Breeding

A
monogamous Monogamy ( ) is a relationship of two individuals in which they form a mutual and exclusive intimate partnership. Having only one partner at any one time, whether for life or serial monogamy, contrasts with various forms of non-monogamy (e.g. ...
species, the male great curassow may build the nest and attract a female's attention to it, though in other cases both members of a pair will build the nest structure. Two eggs are typically laid in a relatively small nest (usually made largely of leaves), each egg measuring and weighing . The young curassow weighs upon hatching; as a half-year-old immature fledgling; and by a year of age, when fully fledged and independent of parental care, will be about three-quarters of their adult weight at .


Diet

Its diet consists mainly of
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
s,
figs The fig is the edible fruit of ''Ficus carica'', a species of tree or shrub in the flowering plant family Moraceae, native to the Mediterranean region, together with western and southern Asia. It has been cultivated since ancient times and i ...
and
arthropods Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
. Small
vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...
s may supplement the diet on occasion, including small
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s (such as
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s). Unlike other cracids, such as guans, they feed largely on fallen fruit rather than pluck fruit directly from the trees. In
Tamaulipas Tamaulipas, officially the Free and Sovereign State of Tamaulipas, is a state in Mexico; one of the 31 states which, along with Mexico City, comprise the 32 federal entities of Mexico. It is divided into 43 municipalities. It is located in nor ...
, it feeds largely on the fruit ''
Spondias mombin ''Spondias mombin'', also known as yellow mombin, hog plum, amra or cajazeira, is a species of tree and flowering plant in the family Anacardiaceae. It is native to the tropical Americas, including the West Indies. The tree was introduced by the ...
''. Elsewhere, it may prefer the red berries of '' Chione'' trees.


Behavior and predators

This species has been noted for its rather aggressive temperament, which has been regularly directed at humans when the birds are held in captivity. Undoubtedly, they have this inclination in order to repel natural predators, from both themselves and their offspring. Known natural predators of this species have included
ocelot The ocelot (''Leopardus pardalis'') is a medium-sized spotted Felidae, wild cat that reaches at the shoulders and weighs between on average. It is native to the southwestern United States, Mexico, Central America, Central and South America, ...
s and ornate hawk-eagles, though chicks and eggs likely have a broader range of predators. When a potential predator is near their offspring, curassows have been noted to engage in a
distraction display Distraction displays, also known as diversionary displays, or paratrepsis are antipredator adaptation, anti-predator behaviors used to attract the attention of an enemy away from something, typically the nest or young, that is being protected by a ...
, feigning injury. When attacking humans, the curassows leap in fluttering flight and scratch about the head, targeting the eyes. Their lifespan in captivity has reached at least 24 years.


Status

Due to ongoing
habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
and overhunting in some areas, the great curassow is evaluated as Vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
of Threatened Species. It is listed on Appendix III of
CITES CITES (shorter acronym for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of inte ...
in
Costa Rica Costa Rica, officially the Republic of Costa Rica, is a country in Central America. It borders Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, and the Pacific Ocean to the southwest, as well as Maritime bo ...
,
Guatemala Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize, to the east by Honduras, and to the southeast by El Salvador. It is hydrologically b ...
, Colombia and
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. It is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Ocean at the Gulf of Fonseca, ...
. Of the smaller
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
''C. r. griscomi'' of Cozumel Island, only a few hundred remain. Its population seems either to have been slowly increasing since the 1980s, or to be fluctuating at a low level; it is vulnerable to
hurricane A tropical cyclone is a rapidly rotating storm system with a low-pressure area, a closed low-level atmospheric circulation, strong winds, and a spiral arrangement of thunderstorms that produce heavy rain and squalls. Depending on its ...
s. This species has proven to produce fertile hybrids with its closest living relative, the blue-billed curassow, and also with the much more distantly related black curassow. In Mexico, there are Unidades de Manejo para la Conservación de la Vida Silvestre anagement Units for the Conservation of Wildlife(UMAs) who are breeding great curassows in captivity.


References


External links

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{{Taxonbar, from=Q572638 great curassow great curassow Birds of Central America Birds of the Yucatán Peninsula Birds of Mexico Birds of Colombia Birds of Ecuador great curassow great curassow Birds of the Sierra Madre Oriental