Cow sharks are a
shark
Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
, the Hexanchidae, characterized by an additional pair or pairs of
gill slit
Gill slits are individual openings to gills, i.e., multiple gill arches, which lack a single outer cover. Such gills are characteristic of cartilaginous fish such as sharks and rays, as well as deep-branching vertebrates such as lampreys. In co ...
s. Its 37 species are placed within the 10 genera: ''
Gladioserratus'', ''
Heptranchias
''Heptranchias'' is a genus of sharks in the family Hexanchidae.
Species
*''Heptranchias perlo'' ( Bonnaterre, 1788) (sharpnose sevengill shark)
*†''Heptranchias ezoensis'' Applegate & Uyeno, 1968
*†''Heptranchias howelli'' Reed, 1946
*� ...
'', ''
Hexanchus
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
'', ''Notidanodon'', ''
Notorynchus
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jordan & ...
'', ''Pachyhexanchus'', ''Paraheptranchias'', ''Pseudonotidanus'', ''
Welcommia'', and ''Weltonia''.
[Compagno, L., Dando, M. and Fowler, S. ''Sharks of the World''. Princeton Field Guides ]
Cow sharks are considered the most primitive of all the sharks, as their skeletons resemble those of ancient extinct forms, with few modern adaptations. Their
excretory and
digestive systems are also unspecialized, suggesting they may resemble those of primitive shark ancestors. A possible hexanchid tooth is known from the Permian of Japan, making the family a possible
extant
Extant is the opposite of the word extinct. It may refer to:
* Extant hereditary titles
* Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English
* Extant taxon, a taxon which is not extinct, ...
survivor of the
Permian-Triassic extinction.
Their most distinctive feature, however, is the presence of a sixth, and, in two genera, a seventh, gill slit, in contrast to the five found in all other sharks.
They range from in adult body length.
Cow sharks are
ovoviviparous
Ovoviviparity, ovovivipary, ovivipary, or aplacental viviparity is a term used as a "bridging" form of reproduction between egg-laying oviparous and live-bearing viviparous reproduction. Ovoviviparous animals possess embryos that develop insi ...
, with the mother retaining the egg cases in her body until they hatch. They feed on relatively large fish of all kinds, including other sharks, as well as on crustaceans and carrion.
The only fossil records of the cow shark consist of mainly only isolated teeth. Although skeletal remains for this species have been found from the Jurassic time period, these have been very rare and have only been found in the "Late Jurassic lithographic limestones of South Germany, Nusplingen, Solnhofen, and late Cretaceous calcareous sediments of Lebanon." Due to these sparse records some scientists conclude that the cow shark is now a more "diverse and numerous species".
Species

The 37 species of cow shark, in 10 genera, are:
* †''
Gladioserratus''
Underwood, Goswami, Prasad, Verma & Flynn, 2011
** †''
Gladioserratus aptiensis''
Pictet, 1864
** †''
Gladioserratus dentatus
''Gladioserratus'' is an extinct genus of cow shark. It contains three species:
*'' Gladioserratus aptiensis'' Pictet, 1865
*'' Gladioserratus magnus'' Underwood, Goswami, Prasad, Verma & Flynn, 2011
*'' Gladioserratus dentatus'' Guinot, Cappe ...
''
Guinot, Cappetta & Adnet, 2014
** †''
Gladioserratus magnus''
Underwood, Goswami, Prasad, Verma & Flynn, 2011
* ''
Heptranchias
''Heptranchias'' is a genus of sharks in the family Hexanchidae.
Species
*''Heptranchias perlo'' ( Bonnaterre, 1788) (sharpnose sevengill shark)
*†''Heptranchias ezoensis'' Applegate & Uyeno, 1968
*†''Heptranchias howelli'' Reed, 1946
*� ...
''
Rafinesque
Constantine Samuel Rafinesque-Schmaltz (; October 22, 1783September 18, 1840) was a French 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ultimat ...
, 1810
** ''
Heptranchias perlo
The sharpnose sevengill shark (''Heptranchias perlo''), also known as one-finned shark, perlon shark, sevengill cow shark, sharpsnouted sevengill or slender sevengill, is a species of shark in the family Hexanchidae, and the only living species ...
''
(Bonnaterre
Abbé Pierre Joseph Bonnaterre (1752, Aveyron – 20 September 1804, Saint-Geniez-d'Olt) was a French zoologist who contributed sections on cetaceans, mammals, birds, reptiles, amphibians
Amphibians are four-limbed and ectothermic ...
, 1788) (sharpnose sevengill shark)
** †''
Heptranchias ezoensis
''Heptranchias'' is a genus of sharks in the family Hexanchidae.
Species
*'' Heptranchias perlo'' ( Bonnaterre, 1788) (sharpnose sevengill shark)
*†'' Heptranchias ezoensis'' Applegate & Uyeno, 1968
*†''Heptranchias howelli'' Reed, 1946 ...
''
Applegate & Uyeno, 1968
** †''
Heptranchias howelli
''Heptranchias howelli'', the sevengill shark, is a nektonic carnivore in the genus '' Heptranchias''. It is an extinct species that ranged from 37.2 to 20.43 Ma.[Heptranchias karagalensis
''Heptranchias'' is a genus of sharks in the family Hexanchidae.
Species
*'' Heptranchias perlo'' ( Bonnaterre, 1788) (sharpnose sevengill shark)
*†'' Heptranchias ezoensis'' Applegate & Uyeno, 1968
*†'' Heptranchias howelli'' Reed, 194 ...]
''
Kozlov in Zhelezko & Kozlov, 1999
** †''
Heptranchias tenuidens
''Heptranchias'' is a genus of sharks in the family Hexanchidae.
Species
*'' Heptranchias perlo'' ( Bonnaterre, 1788) (sharpnose sevengill shark)
*†'' Heptranchias ezoensis'' Applegate & Uyeno, 1968
*†'' Heptranchias howelli'' Reed, 194 ...
''
Leriche, 1938
* ''
Hexanchus
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
''
Rafinesque, 1810
** ''
Hexanchus griseus
The bluntnose sixgill shark (''Hexanchus griseus''), often simply called the cow shark, is the largest hexanchoid shark, growing to in length. It is found in tropical and temperate waters worldwide and its diet is widely varied by region.
The ...
''
(Bonnaterre, 1788) (bluntnose sixgill shark)
** ''
Hexanchus nakamurai
The bigeyed sixgill shark (''Hexanchus nakamurai'') is a cow shark of the family Hexanchidae. Its dorsal surface has a brownish-gray color, and is sharply separated from the light coloring of its ventral surface. The eyes are a fluorescent gre ...
''
Teng Teng may refer to:
* Teng (surname) (滕), a Chinese surname
* Teng (state), an ancient Chinese state
* Teng (mythology), a flying dragon in Chinese mythology
*Teng County
Teng County or Tengxian (; za, Dwngz Yen) is a county of eastern Guangx ...
, 1962 (bigeyed sixgill shark)
** ''
Hexanchus vitulus
The Atlantic sixgill shark (''Hexanchus vitulus'') is a rare species of hexanchid shark found in the Atlantic Ocean. It was formerly described as its own species, but was synonymised with the bigeye sixgill shark
The bigeyed sixgill shark ...
''
Daly-Engel, 2018 (atlantic sixgill shark)
** †''
Hexanchus agassizi''
Cappetta, 1976
** †''
Hexanchus andersoni
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
''
Jordan, 1907
** †''
Hexanchus casieri
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
''
Kozlov, 1999
** †''
Hexanchus collinsonae
''Hexanchus collinsonae'' (named after M. E. Collinson) is a species of prehistoric shark that was found in the Eocene London Clay beds. It is a member of the family
Family (from la, familia) is a Social group, group of people related eit ...
''
Ward, 1979
** †''
Hexanchus gracilis
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
''
Davis, 1887
** †''
Hexanchus hookeri
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
''
Ward, 1979
** †''
Hexanchus microdon
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
''
Agassiz, 1843
** †''
Hexanchus tusbairicus
The sixgill sharks are a genus, ''Hexanchus'', of deepwater sharks in the family Hexanchidae. These sharks are characterized by a broad, pointed head, six pairs of gill slits, comb-like, yellow lower teeth, and a long tail. The largest species ca ...
''
Kozlov in Zhelezko & Kozlov, 1999
* †''
Notidanodon''
Cappetta, 1975
** †''
Notidanodon brotzeni
''Notidanodon'' is an extinct genus of cow shark. Fossils ascribed to this genus are known from the Jurassic, Cretaceous, and Paleogene periods. The genus is known from every continent including Antarctica.
Species
The earliest occurrence ...
''
Siverson, 1995
** †''
Notidanodon dentatus
''Notidanodon'' is an extinct genus of cow shark. Fossils ascribed to this genus are known from the Jurassic, Cretaceous, and Paleogene periods. The genus is known from every continent including Antarctica.
Species
The earliest occurrence ...
''
Woodward, 1886
** †''
Notidanodon lanceolatus
''Notidanodon'' is an extinct genus of cow shark. Fossils ascribed to this genus are known from the Jurassic, Cretaceous, and Paleogene periods. The genus is known from every continent including Antarctica.
Species
The earliest occurrences of ...
''
Woodward, 1886
** †''
Notidanodon loozi
''Notidanodon'' is an extinct genus of cow shark. Fossils ascribed to this genus are known from the Jurassic, Cretaceous, and Paleogene periods. The genus is known from every continent including Antarctica.
Species
The earliest occurrence ...
''
Vincent, 1876
** †''
Notidanodon pectinatus
''Notidanodon'' is an extinct genus of cow shark. Fossils ascribed to this genus are known from the Jurassic, Cretaceous, and Paleogene periods. The genus is known from every continent including Antarctica.
Species
The earliest occurrence ...
''
Agassiz, 1843
* ''
Notorynchus
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jordan & ...
''
Ayres, 1855
** ''
Notorynchus cepedianus''
( Péron, 1807) (broadnose sevengill shark)
** †''
Notorynchus borealus
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jordan & ...
''
Jordan & Hannibal, 1923
** †''
Notorynchus kempi
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jordan & ...
''
Ward, 1979
** †''
Notorynchus lawleyi
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jordan & ...
''
Cigala Fulgosi, 1983
** †''
Notorynchus primigenius
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jordan & ...
''
Agassiz, 1843
** †''
Notorynchus serratissimus
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jord ...
''
Agassiz, 1843
** †''
Notorynchus subrecurvus
''Notorynchus'' is a genus of deepwater sharks in the family Hexanchidae. There is one extant species.
Extant species
* Broadnose sevengill shark (''Notorynchus cepedianus'') Péron, 1807
Extinct species
* '' Notorynchus borealus'' Jordan & ...
''
Oppenheimer, 1907
* †''
Pachyhexanchus''
Cappetta, 1990
** †''
Pachyhexanchus pockrandti''
Ward & Thies, 1987
* †''
Paraheptranchias''
Pfeil, 1981
** †''
Paraheptranchias repens''
Probst, 1879
* †''
Pseudonotidanus''
Underwood & Ward, 2004
** †''
Pseudonotidanus semirugosus''
Underwood & Ward, 2004
* †''
Welcommia''
Klug & Kriwet, 2010
** †''
Welcommia bodeuri''
Cappetta, 1990
** †''
Welcommia cappettai
''Welcommia'' is an extinct genus of shark. It contains only two species - ''Welcommia bodeuri'' and ''Welcommia cappettai'', described from the middle Oxfordian of Germany by Stefanie Klug and Jürgen Kriwet, in 2010.Stefanie Klug and Jürgen ...
''
Klug & Kriwet, 2010
* †''
Weltonia''
Ward, 1979
** †''
Weltonia ancistrodon''
Arambourg, 1952
** †''
Weltonia burnhamensis''
Ward, 1979
References
External links
ReefQuest Centre: ''Order Hexanchiformes: Cow Sharks''
{{DEFAULTSORT:Cow Shark
Hexanchiformes
Extant Late Jurassic first appearances
Taxa named by John Edward Gray