Corsia Ornata
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''Corsia ornata'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Corsia'' of the small
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Corsiaceae, part of the
monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are flowering plants whose seeds contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. A monocot taxon has been in use for several decades, but ...
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Liliales Liliales is an order (biology), order of monocotyledonous flowering plants in the Angiosperm Phylogeny Group and Angiosperm Phylogeny Web List of systems of plant classification, system, within the lilioid monocots. This order of necessity includ ...
. They are saprophytes (
Myco-heterotrophs Myco-heterotrophy (from Greek , , and ) is a symbiotic relationship between certain kinds of plants and fungi, in which the plant gets all or part of its food from parasitism upon fungi rather than from photosynthesis. A myco-heterotroph i ...
), lacking the ability to photosynthesise, being dependent on other organisms for their nutrition. The plant lives underground, sending up purplish stems above ground in order to flower. The leaves are reduced to scales. One of the six petal-like
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of ve ...
named the labellum (a little lip), is specialised, being enlarged and hanging protectively over the reproductive organs. It was discovered in New Guinea in 1875, but has since been sighted in Queensland, Australia.


Description

The Corsiaceae lack
chlorophyll Chlorophyll is any of several related green pigments found in cyanobacteria and in the chloroplasts of algae and plants. Its name is derived from the Greek words (, "pale green") and (, "leaf"). Chlorophyll allows plants to absorb energy ...
and hence the ability to photosynthesise, instead being
mycoheterotrophic Myco-heterotrophy (from Greek , , and ) is a symbiotic relationship between certain kinds of plants and fungi, in which the plant gets all or part of its food from parasitism upon fungi rather than from photosynthesis. A myco-heterotroph i ...
(deriving nutrition as parasites on fungi). ''C. ornata'', although perennial, only appears above ground when flowering, arising from short creeping
rhizomes In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
, reaching up to 25 cm in height. From the rhizomes, arise long, cylindrical and finely corrugated, unbranched and upright growing stems that are
terete Terete is a term in botany used to describe a cross section (geometry), cross section that is circular, or like a distorted circle, with a single surface wrapping around it.Lichen Vocabulary, Lichens of North America Information, Sylvia and S ...
(almost circular in cross section) and narrowly ribbed. The above ground portion of the plant is a purplish to purplish-red in color.
Leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are reduced to acute (sharply pointed) sheath-shaped scales 1–2 cm in length, arranged alternately on the stem, with 3–5
nerves A nerve is an enclosed, cable-like bundle of nerve fibers (called axons). Nerves have historically been considered the basic units of the peripheral nervous system. A nerve provides a common pathway for the electrochemical nerve impulses called ...
and similar
bracts In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also loo ...
. The
pedicels In botany, a pedicel is a stem that attaches a single flower to the inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branch ...
are
glabrous Glabrousness () is the technical term for a lack of hair, down, setae, trichomes, or other such covering. A glabrous surface may be a natural characteristic of all or part of a plant or animal, or be due to loss because of a physical condition, ...
and 2.5–4 cm long. The upright individual flowers are terminal and stand on flower stems that are 2.5–4 cm long. Of the six
tepals A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of ve ...
(in two
whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). In nature File:Photograph and axial plane floral diagram ...
), five are linear, obtuse and pale yellow in colour, 11–13  millimeters long, one-nerved and hairless. The sixth, outer tepal, called the labellum, is either light yellow to light purple with a darker purple veins nerve or purplish brown. It is greatly enlarged (1.2–1.8 cm long, 1–1.6 cm wide) and cordate (heart-shaped) and obtuse, with a cordate base. The labellum initially surrounds the flower bud and, after opening, protects the other flower organs. It has a basal callus that is white, broadly shell-shaped, 2–3.5 mm long and around 2.5 mm wide, with a tip that is rounded or slightly acuminate, finely papillate at the margins with 8 or 9 lateral nerves that are variously branched and 16–18 short
lamellae Lamella (: lamellae) means a small plate or flake in Latin, and in English may refer to: Biology * Lamella (mycology), a papery rib beneath a mushroom cap * Lamella (botany) * Lamella (surface anatomy), a plate-like structure in an animal * Lame ...
radiating from the basal callus that are distinctly
pilose Trichomes (; ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...
. At the base, the labellum overhangs the reproductive organs umbrella-like, the 1 mm long
gynostemium The column, or technically the gynostemium, is a reproductive structure that can be found in several plant families: Aristolochiaceae, Orchidaceae, and Stylidiaceae. It is derived from the fusion of both male and female parts (stamens and pistil ...
(fused
stamens The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filamen ...
and
pistil Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
). The 1.5 mm filaments are dark purple, the anthers dark purple to pink, 0.5 by 1 mm. The
style Style, or styles may refer to: Film and television * ''Style'' (2001 film), a Hindi film starring Sharman Joshi, Riya Sen, Sahil Khan and Shilpi Mudgal * ''Style'' (2002 film), a Tamil drama film * ''Style'' (2004 film), a Burmese film * '' ...
is about 1.5 mm in length and the capsule 2.5–3.5 cm.


Taxonomy

''Corsia ornata'' was first described by
Odoardo Beccari Odoardo Beccari (16 November 1843 – 25 October 1920) was an Italian botanist famous for his discoveries in Indonesia, New Guinea, and Australia. He has been called the greatest botanist to ever study Malesia. Life Youth and education (18 ...
in February 1875, and therefore bears his name (Becc.) as the
botanical authority In botanical nomenclature, author citation is the way of citing the person or group of people who validly published a botanical name, i.e. who first published the name while fulfilling the formal requirements as specified by the ''International Cod ...
. It was the first
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
to be discovered and is therefore considered its
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
. It is classified in the
section Section, Sectioning, or Sectioned may refer to: Arts, entertainment and media * Section (music), a complete, but not independent, musical idea * Section (typography), a subdivision, especially of a chapter, in books and documents ** Section sig ...
''Sessilis'' (named for their sessile labellum), the larger of the two sections of that genus, as one of 19 species.


Distribution and habitat

''Corsia ornata'' is found at a number of widely spaced locations in the western part of New Guinea, particularly on
Bird's Head Peninsula The Bird's Head Peninsula ( Indonesian: , , meaning Bird's Head in Indonesian and Dutch) or Doberai Peninsula (''Semenanjung Doberai'') is a large peninsula that makes up the northwest portion of the island of New Guinea, comprising the Indones ...
. One sighting in Queensland, Australia, makes it the only species found outside of New Guinea. ''Corsia ornata'' occurs in forests, between 400 and 2,100 m, in humus rich soils.


References


Bibliography

;Books * * * ;Articles * * ;Websites * * *


External links

{{Taxonbar, from= Q1135465 Corsiaceae Parasitic plants Parasites of fungi Taxa named by Odoardo Beccari