The Corispermoideae are a subfamily of the

Amaranthaceae Amaranthaceae ( ) is a family of flowering plants commonly known as the amaranth family, in reference to its type genus '' Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, maki ...

, formerly in family Chenopodiaceae.

Description

The species of the subfamily Corispermoideae are all

annual plant An annual plant is a plant that completes its life cycle, from germination to the production of seeds, within one growing season, and then dies. Globally, 6% of all plant species and 15% of herbaceous plants (excluding trees and shrubs) are ...

s. Leaves are mostly alternate, sessile or petiole-like attenuate, laminate, scleromorphic. Typical are branched (dendritic)

trichome Trichomes (; ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...

s (except in '' Anthochlamys'') on young plant parts. The flowers are arranged in simple, compact (sometimes globular) partial

inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...

s, or in spikes. Bracteoles are missing. The

perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepal ...

consists of 1-5 white, membranaceous

tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...

s (missing in some '' Corispermum'' species) without

vascular bundle A vascular bundle is a part of the transport system in vascular plants. The transport itself happens in the stem, which exists in two forms: xylem and phloem. Both these tissues are present in a vascular bundle, which in addition will incl ...

s, not persistent. The

pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...

grains of '' Agriophyllum'' and '' Corispermum'' are of the "Chenopodium type", of '' Anthochlamys'' of the "Anthochlamys type". The fruits possess supporting tissue consisting of macro

sclereid Sclereids are a reduced form of sclerenchyma cells with highly thickened, lignified cellular walls that form small bundles of durable layers of tissue in most plants.Evert, Ray F; Eichhorn, Susan E. Esau's Plant Anatomy: Meristems, Cells, and T ...

s. The seeds have a vertical embryo and copious

perisperm In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fema ...

.Alexander P. Sukhorukov: ''Fruit anatomy and its taxonomic significance in Corispermum (Corispermoideae, Chenopodiaceae).'' – Willdenowia 37, 2007, , p.63-87, ,
pdf

Distribution

The subfamily Corispermoideae is distributed in Asia, Europe and North America.

Photosynthesis pathway

All species studied show non-Kranz corispermoid leaf anatomy and C₃ photosynthesis.

Taxonomy

The tribe Corispermeae was published in 1840 by

Alfred Moquin-Tandon Christian Horace Benedict Alfred Moquin-Tandon (7 May 1804 – 15 April 1863) was a French Natural history, naturalist and doctor. Moquin-Tandon was professor of zoology at Marseille from 1829 until 1833, when he was appointed professor of botany ...

(in ''Chenopodearum Monographica Enumeratio'', Loss, Paris, S. 182). Oskar Eberhard Ulbrich raised it to subfamily level named Corispermoideae in 1934 (in ''Chenopodiaceae'', S. 379–584 in

Adolf Engler Heinrich Gustav Adolf Engler (25 March 1844 – 10 October 1930) was a German botanist. He is notable for his work on plant taxonomy and phytogeography, such as ''Die natürlichen Pflanzenfamilien'' (''The Natural Plant Families''), edited with K ...

Karl Anton Eugen Prantl Karl Anton Eugen Prantl (10 September 1849 – 24 February 1893), also known as Carl Anton Eugen Prantl, was a German botanist. Prantl was born in Munich, Kingdom of Bavaria, and studied in Munich. In 1870, he graduated with the dissertation ''D ...

(Edt.): ''Die natürlichen Pflanzenfamilien'', Band 16 c, Engelmann, Leipzig). Molecular data support the

monophyly In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent comm ...

of the subfamily. It contains only one tribe: * Tribus Corispermeae, with 3 genera: ** '' Agriophyllum'' M.Bieb., with 6 species in Asian regions with arid climate ** '' Anthochlamys'' Fenzl., with 2 species in Asian regions with arid climate ** '' Corispermum'' L., with at least 65 species in extratropical regions of Eurasia and North AmericaJuan Juan Xue & Ming Li Zhang: ''Monophyly and infrageneric variation of Corispermum L. (Chenopodiaceae), evidence from sequence data psbB-psbH, rbcL and ITS'', In: ''Journal of Arid Land'', Volume 3, Issue 4, 2011, p. 240−253.
Fulltext-PDF.
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References

External links

Chenopodiaceae, p. 72
un
p. 73
Georg Volkens: ''Cyclolobaeae-Corispermeae'' i
Adolf Engler & Karl Anton Eugen Prantl: ''Die Natürlichen Pflanzenfamilien'', 3., 1893.
{{Taxonbar, from=Q15962912 Amaranthaceae Caryophyllales subfamilies Taxa named by Oskar Eberhard Ulbrich