Coptoclavella
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''Coptoclavella'' is an extinct genus of beetles in the family
Coptoclavidae Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Adephaga, Dytiscoidea, which contains living ...
. There are about seven described species in ''Coptoclavella''.


Species

These seven species belong to the genus ''Coptoclavella'': * † ''Coptoclavella elegans'' Ponomarenko, 1980Mogotuin Formation, Mongolia,
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
* † ''Coptoclavella inexpecta'' Soriano ''et al.'', 2007
La Pedrera de Rúbies Formation The La Pedrera de Rúbies Formation, also called as La Pedrera de Meià is an Early Cretaceous ( late Berriasian to early Barremian geologic formation in Catalonia, Spain. The formation crops out in the area of the Montsec in the Organyà Bas ...
, Spain,
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
* † ''Coptoclavella jurassica'' Ponomarenko, 2014Sharteg, Mongolia,
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
* † ''Coptoclavella minor'' Ponomarenko, 1980Takshin Formation,
Kalgan Formation Zhangjiakou (), also known as Kalgan and by several other names, is a prefecture-level city in northwestern Hebei province in Northern China, bordering Beijing to the southeast, Inner Mongolia to the north and west, and Shanxi to the southwest ...
Russia,
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
;
Daya Formation Daya may refer to: Religion * Daya (Sikhism), the concept of compassion in Sikhism * Daya (virtue), the concept of compassion in Hinduism Media and music * ''Daya'' (film), a 1998 Malayalam film * ''Daya'' (EP), a 2015 recording by the America ...
, Russia,
Hauterivian The Hauterivian is, in the geologic timescale, an age in the Early Cretaceous Epoch or a stage in the Lower Cretaceous Series. It spans the time between 132.6 ± 2 Ma and 125.77 (million years ago). The Hauterivian is preceded by the Valangi ...
; Mogotuin Formation, Mongolia, Aptian; Turga Formation, Russia, Aptian * † ''Coptoclavella purbeckensis'' Ponomarenko ''et al.'', 2005
Durlston Formation The Durlston Formation is a geologic formation in England. Particularly in the Isle of Purbeck. It preserves fossils dating back to the Berriasian stage of the Lower Cretaceous. Vertebrate paleobiota Crocodyliformes See also * List of ...
, United Kingdom,
Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...
* † ''Coptoclavella striata'' Ponomarenko, 1986Gurvan-Eren Formation, Mongolia, Aptian * † ''Coptoclavella vittata'' Ponomarenko, 1986 – Gurvan-Eren Formation, Mongolia, Aptian


References

Coptoclavidae Adephaga genera Fossil beetle genera {{Adephaga-stub