''Copelatus'' is a large genus of small

diving beetle The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive", are the predaceous diving beetles, a Family (biology), family of water beetles. They occur in virtually any freshwater habitat around the world, bu ...

s. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in color, many species of Copelatus possessing visible longitudinal furrows down the dorsal side of the wings of both sexes.

Systematics

The genus ''Copelatus'' is divided into several subgenera (''Colepatus'', ''Papuadytes'' etc.), some of which are sometimes treated as separate genera. It may be

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

with respect to the smaller Copelatinae genera '' Lacconectus'' and '' Aglymbus''.Balke,M et al., MtDNA phylogeny and biogeography of Copelatinae, a highly diverse group of tropical diving beetles (Dytiscidae). Molecular Phylogenetics and Evolution Vol. 32, Issue 3, 2004 The species include:

Species

* '' Copelatus abonnenci'' Guignot, 1939 * '' Copelatus acamas'' Guignot, 1955 * '' Copelatus adelbert'' Megna, Atthakor, Manaono, Hendrich & Balke, 2017 * '' Copelatus advena'' Sharp, 1882 * '' Copelatus aemulus'' Bilardo & Rocchi, 1995 * '' Copelatus aequatorius'' Régimbart, 1899 * '' Copelatus aethiopicus'' Régimbart, 1906 * '' Copelatus agaroensis'' Balke, 2008 * '' Copelatus agrias'' Guignot, 1954 * '' Copelatus alternatus'' Sharp, 1882 * '' Copelatus amaroides'' Guignot, 1952 * ''

Copelatus amatolensis ''Copelatus amatolensis'' is a species of diving beetle. It is part of the genus ''Copelatus'' of the subfamily Copelatinae in the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to di ...

'' Omer-Cooper, 1965 * '' Copelatus amazonicus'' Régimbart, 1889 * '' Copelatus ambiguus'' Bertrand & Legros, 1975 * ''

Copelatus amphibius ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * ''

Copelatus anastomosans ''Copelatus anastomosans'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to d ...

'' Guignot, 1952 * ''

Copelatus andobonicus ''Copelatus andobonicus'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to di ...

'' Guignot, 1960 * ''

Copelatus andreinii ''Copelatus andreinii'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive ...

'' Régimbart, 1905 * ''

Copelatus andrewesi ''Copelatus andrewesi'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive ...

'' J. Balfour-Browne, 1939 * ''

Copelatus angolensis ''Copelatus angolensis'' is a species of diving beetle. It is part of the genus ''Copelatus'' of the subfamily Copelatinae in the family Dytiscidae. It was described by Peschet in 1924.Copelatus angouei ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2011 * ''

Copelatus angustus ''Copelatus angustus'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive" ...

'' Gschwendtner, 1932 * ''

Copelatus ankaratra ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * ''

Copelatus annobomensis ''Copelatus annobomensis'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to d ...

'' J. Balfour-Browne, 1939 * ''

Copelatus anthracinus ''Copelatus anthracinus'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to di ...

'' Régimbart, 1895 * ''

Copelatus antoniorum ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek & Brancucci, 2011 *†'' Copelatus aphroditae'' Balke, 2003 * ''

Copelatus apicinotatus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2018 * ''

Copelatus assalyi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2010 * '' Copelatus assimilis'' Régimbart, 1895 * '' Copelatus ateles'' Guignot, 1955 * '' Copelatus atrosulcatus'' Régimbart, 1906 * ''

Copelatus babyrousa ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Hendrich & Balke, 2018 * '' Copelatus bacchusi'' Wewalka, 1981 * '' Copelatus bacillifer'' Guignot, 1955 * '' Copelatus baculiformis'' Guignot, 1955 * '' Copelatus badeni'' Sharp, 1882 * '' Copelatus bakewelli'' J. Balfour-Browne, 1939 * '' Copelatus baoulicus'' Bilardo & Pederzani, 1978 * '' Copelatus bapounouensis'' Bilardo & Rocchi, 2019 * '' Copelatus baranensis'' Hájek, Shaverdo, Hendrich & Balke, 2021 * '' Copelatus barbouri'' Young, 1942 * '' Copelatus basifasciatus'' Régimbart, 1895 * '' Copelatus basilewskyi'' Bilardo & Pederzani, 1979 * '' Copelatus bechynei'' Guignot, 1953 * '' Copelatus befasicus'' Guignot, 1956 * '' Copelatus bengalensis'' Guignot, 1955 * '' Copelatus bertrandi'' Nilsson, Bilardo & Rocchi, 1996 * ''

Copelatus betampona ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * ''

Copelatus bezdeki ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Sheth, Ghate & Hájek, 2018 * '' Copelatus bibulus'' Guignot, 1948 * '' Copelatus biformis'' Sharp, 1882 * '' Copelatus bimaculatus'' (Resende & Vanin, 1991) * '' Copelatus binaghii'' Bilardo & Pederzani, 1978 * ''

Copelatus biolleyi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Guignot, 1952 * ''

Copelatus biroi ''Copelatus biroi'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by Félix Guignot, Guignot in 1956.Copelatus birougouensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2018 * ''

Copelatus biseriatus ''Copelatus biseriatus'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by J. Balfour-Browne in 1939.Copelatus biswasi'' Mukherjee & Sengupta, 1986 * '' Copelatus blancasi'' Guignot, 1958 * '' Copelatus blatchleyi'' Young, 1953 NA * '' Copelatus bolivianus'' Guignot, 1957 * '' Copelatus bombycinus'' Guignot, 1952 * '' Copelatus bonvouloiri'' Sharp, 1882 * '' Copelatus bottegoi'' Régimbart, 1895 * '' Copelatus bougainvillensis'' Hájek, Shaverdo, Hendrich & Balke, 2021 * '' Copelatus boukali'' Hendrich & Balke, 1998 * '' Copelatus brancuccii'' Rocchi, 1979 * '' Copelatus brasiliensis'' Zimmermann, 1921 * ''

Copelatus brendelli ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Hendrich & Balke, 2018 * '' Copelatus brevicornis'' (Sharp, 1882) * '' Copelatus brevistrigatus'' Guignot, 1959 * '' Copelatus brivioi'' Rocchi, 1976 * '' Copelatus bromeliarum'' (Scott, 1912) * '' Copelatus brullei'' Aubé, 1838 * '' Copelatus brunneus'' J. Balfour-Browne, 1939 * '' Copelatus buqueti'' Aubé, 1838 * '' Copelatus burgeoni'' Gschwendtner, 1930 * '' Copelatus caelatipennis'' Aubé, 1838 * '' Copelatus caelatus'' Guignot, 1952 * '' Copelatus caffer'' J. Balfour-Browne, 1939 * ''

Copelatus calaquei ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2008 * '' Copelatus camerunensis'' Guignot, 1941 * '' Copelatus capensis'' Sharp, 1882 * '' Copelatus carayoni'' Legros, 1949 * '' Copelatus carinatus'' Sharp, 1882 * '' Copelatus celinoides'' Guignot, 1952 * '' Copelatus cessaima'' Caetano, Bená & Vanin, 2013 * ''

Copelatus chevrolati ''Copelatus chevrolati'' is a species of Dytiscidae, diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by Aubé in 1838.Copelatus chibcha'' Guignot, 1952 * ''

Copelatus chinensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Régimbart, 1899 * '' Copelatus chipiriricus'' Guignot, 1957 * '' Copelatus chloroticus'' Régimbart, 1899 * '' Copelatus cinnamomeus'' Régimbart, 1895 * '' Copelatus clarki'' Sharp, 1882 * '' Copelatus collarti'' Gschwendtner, 1932 * '' Copelatus compertus'' Guignot, 1956 * '' Copelatus concii'' Bilardo, 1982 * '' Copelatus concolor'' Sharp, 1882 * '' Copelatus concolorans'' J. Balfour-Browne, 1939 * '' Copelatus confinis'' Bilardo & Rocchi, 1999 * '' Copelatus congo'' Gschwendtner, 1938 * '' Copelatus consimilis'' Bilardo & Rocchi, 2002 * '' Copelatus consors'' Sharp, 1882 *†''

Copelatus convexus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' (Förster, 1891) * '' Copelatus cooperae'' Bilardo & Pederzani, 1972 * '' Copelatus cordovai'' Megna & Epler, 2012 * '' Copelatus cordylinoides'' J. Balfour-Browne, 1938 * '' Copelatus coxalis'' Sharp, 1882 * '' Copelatus crassus'' Régimbart, 1895 * '' Copelatus cryptarchoides'' Régimbart, 1899 * '' Copelatus cubaensis'' Schaeffer, 1908 * '' Copelatus curtistriatus'' Bilardo & Rocchi, 1995 * ''

Copelatus curvispinis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2008 * ''

Copelatus cynthiae ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2008 * '' Copelatus daemeli'' Sharp, 1882 * '' Copelatus danyi'' Megna & Epler, 2012 * '' Copelatus darlingtoni'' Young, 1942 * ''

Copelatus davidi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Wewalka, 2017 * '' Copelatus debilis'' Sharp, 1882 * ''

Copelatus deccanensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Sheth, Ghate & Hájek, 2018 * ''

Copelatus decellei ''Copelatus decellei'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive" ...

'' Bilardo, 1982 * '' Copelatus decemsulcatus'' Régimbart, 1895 * '' Copelatus deceptor'' Bilardo & Rocchi, 1995 * '' Copelatus dentatipenis'' Jiang, Hájek & Jia, 2022 * '' Copelatus depressus'' Sharp, 1882 * '' Copelatus descarpentriesi'' Bertrand & Legros, 1975 * '' Copelatus diffisus'' Guignot, 1939 * '' Copelatus dimorphus'' Sharp, 1882 * '' Copelatus distinctus'' Aubé, 1838 * '' Copelatus distinguendus'' Régimbart, 1903 * ''

Copelatus diversistriatus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Jiang, Hájek & Jia, 2022 * ''

Copelatus djiboutensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Wewalka & Jäch, 2017 * '' Copelatus dolosus'' Guignot, 1956 * '' Copelatus doriae'' Sharp, 1882 * ''

Copelatus doudouensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2015 * '' Copelatus duodecimstriatus'' Aubé, 1838 * '' Copelatus duponti'' Aubé, 1838 * '' Copelatus efoutensis'' Bilardo & Rocchi, 1995 * '' Copelatus ejactus'' Omer-Cooper, 1965 * '' Copelatus ellai'' Bilardo & Rocchi, 1995 * '' Copelatus elutus'' Guignot, 1958 * '' Copelatus enganensis'' Guignot, 1940 * '' Copelatus epactus'' Guignot, 1948 * '' Copelatus erichsonii'' Guérin-Méneville, 1847 * '' Copelatus esteriensis'' Bilardo & Pederzani, 1978 * '' Copelatus eucritus'' Guignot, 1952 * '' Copelatus evanidus'' Bilardo & Rocchi, 1995 * '' Copelatus exaratus'' Sharp, 1882 * '' Copelatus externus'' Kirsch, 1873 * ''

Copelatus falciformis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Pederzani & Schizzerotto, 2017 * '' Copelatus fasciatus'' Bilardo & Rocchi, 1995 * ''

Copelatus fastidiosus ''Copelatus fastidiosus'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by Félix Guignot Félix Guignot was a French physician and entomologist, b ...

'' Guignot, 1959 * '' Copelatus feae'' Régimbart, 1888 * ''

Copelatus felicis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Jiang & Jia, 2022 * '' Copelatus fernandensis'' Guignot, 1952 * '' Copelatus ferruginicollis'' Régimbart, 1895 * '' Copelatus ferus'' Guignot, 1953 * '' Copelatus festae'' Régimbart, 1899 * '' Copelatus fidschiensis'' Zimmermann, 1928 * '' Copelatus fijiensis'' Guignot, 1955 * '' Copelatus filiformis'' Sharp, 1882 * ''

Copelatus fizpaci ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2013 * '' Copelatus flavicans'' Guignot, 1952 * '' Copelatus flavidus'' Régimbart, 1895 * '' Copelatus fluviaticus'' Guignot, 1957 * '' Copelatus fontanus'' J. Balfour-Browne, 1950 *†'' Copelatus fossilis'' Říha, 1974 * '' Copelatus fractistriatus'' Bilardo & Rocchi, 1995 * '' Copelatus freudei'' Guignot, 1955 * '' Copelatus fryi'' J. Balfour-Browne, 1939 * '' Copelatus fulviceps'' J. Balfour-Browne, 1938 * '' Copelatus fuscipennis'' Sharp, 1882 * '' Copelatus fuscomaculatus'' Guignot, 1952 * '' Copelatus gabonicus'' Bilardo & Pederzani, 1978 * '' Copelatus galapagoensis'' G.R. Waterhouse, 1845 * '' Copelatus garambanus'' Guignot, 1955 * '' Copelatus gardineri'' Scott, 1912 * '' Copelatus geayi'' Régimbart, 1904 * '' Copelatus geniculatus'' Sharp, 1882 * '' Copelatus gentilis'' Sharp, 1882 * ''

Copelatus gestroi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' (Sharp, 1882) * ''

Copelatus gigas ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2019 * '' Copelatus glyphicus'' (Say, 1823) NA * '' Copelatus griffinii'' Régimbart, 1899 * '' Copelatus gschwendtneri'' Guignot, 1939 * '' Copelatus guadelupensis'' Legros, 1948 * ''

Copelatus guerini ''Copelatus guerini'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by Aubé in 1838. References guerini Guerini is an Italian surname. Notable pe ...

'' Aubé, 1838 * '' Copelatus guineensis'' Guignot, 1955 * '' Copelatus haemorrhoidalis'' Régimbart, 1883 * '' Copelatus hararensis'' Guignot, 1952 * '' Copelatus hardenbergi'' J. Balfour-Browne, 1950 * '' Copelatus hebeter'' Guignot, 1953 * '' Copelatus heterogynus'' Régimbart, 1899 * ''

Copelatus hydroporoides ''Copelatus hydroporoides'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to ...

'' (Murray, 1859) * '' Copelatus ibrahimi'' Zalat, Saleh, Angus & Kaschef, 2000 * ''

Copelatus ignacioi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Hendrich & Balke, 2022 * '' Copelatus iguelaensis'' Bilardo & Rocchi, 2002 * '' Copelatus ilybioides'' Régimbart, 1895 * '' Copelatus imitator'' Bilardo & Rocchi, 2002 * '' Copelatus inaequalis'' Sharp, 1882 * '' Copelatus incognitus'' Sharp, 1882 * '' Copelatus indicus'' Sharp, 1882 * '' Copelatus inopinatus'' Bilardo & Rocchi, 1995 * '' Copelatus inornatus'' Sharp, 1882 * '' Copelatus insidiosus'' Bilardo & Rocchi, 1995 * '' Copelatus insolitus'' Chevrolat, 1863 * '' Copelatus instabilis'' Régimbart, 1897 * '' Copelatus instriatus'' Zimmermann, 1921 * '' Copelatus insuetus'' Guignot, 1941 * '' Copelatus insulanus'' Guignot, 1939 * '' Copelatus integer'' Sharp, 1882 * '' Copelatus internus'' Régimbart, 1904 * '' Copelatus interstriatus'' J. Balfour-Browne, 1939 * ''

Copelatus inuber ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Guignot, 1952 * '' Copelatus ipiformis'' Régimbart, 1895 * ''

Copelatus irinus ''Copelatus irinus'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive", ...

'' Régimbart, 1899 * '' Copelatus irregularis'' W.J. Macleay, 1871 * '' Copelatus ischius'' Guignot, 1956 * '' Copelatus jactator'' Guignot, 1955 * ''

Copelatus jaechi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2016 * '' Copelatus jamaicensis'' Young, 1942 * '' Copelatus japonicus'' Sharp, 1884 * '' Copelatus jarrigei'' Legros, 1954 * '' Copelatus javanus'' Régimbart, 1883 * '' Copelatus jocosus'' Guignot, 1955 * '' Copelatus johannis'' Guignot, 1954 * '' Copelatus kalaharii'' Gschwendtner, 1935 * '' Copelatus kammuriensis'' Tamu & Tsukamoto, 1955 * '' Copelatus kaszabi'' Guignot, 1956 * ''

Copelatus kely ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * '' Copelatus kietensis'' Hájek, Shaverdo, Hendrich & Balke, 2021 * '' Copelatus kilimandjarensis'' Bilardo & Pederzani, 1972 * '' Copelatus kindianus'' Guignot, 1954 * '' Copelatus kongouensis'' Bilardo & Rocchi, 1999 * '' Copelatus koreanus'' T. Mori, 1932 * '' Copelatus laccophilinus'' Sharp, 1882 * '' Copelatus laeticulus'' Sharp, 1882 * ''

Copelatus laevipennis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Shaverdo, Hendrich & Balke, 2021 * '' Copelatus lamottei'' Legros, 1954 * '' Copelatus lanzai'' Bilardo & Rocchi, 1995 * '' Copelatus laraensis'' Bilardo & Rocchi, 1995 * '' Copelatus lasckonyi'' Bilardo & Rocchi, 1995 * '' Copelatus latens'' Guignot, 1952 * '' Copelatus laticollis'' Régimbart, 1899 * '' Copelatus latifasciatus'' Bilardo & Rocchi, 1999 * '' Copelatus latipes'' Sharp, 1882 * '' Copelatus latus'' J. Balfour-Browne, 1950 * '' Copelatus leonardii'' Bilardo & Rocchi, 1999 * '' Copelatus leonensis'' Legros, 1954 * '' Copelatus lepersonneae'' Gschwendtner, 1943 * '' Copelatus lignosus'' Guignot, 1955 * '' Copelatus lineatipennis'' Guignot, 1955 * '' Copelatus lineatus'' (Guérin-Méneville, 1838) * '' Copelatus longicornis'' Sharp, 1882 * '' Copelatus lootensi'' Guignot, 1955 * ''

Copelatus louayensis ''Copelatus louayensis'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to div ...

'' Bilardo & Rocchi, 2004 * '' Copelatus luctuosus'' Guignot, 1939 * '' Copelatus luridescens'' Régimbart, 1889 * '' Copelatus luteocinctus'' Guignot, 1955 * '' Copelatus luteomaculatus'' Guignot, 1956 * '' Copelatus luzonicus'' Guignot, 1952 * '' Copelatus macellus'' Guignot, 1950 * '' Copelatus madoni'' Guignot, 1952 * '' Copelatus mahajanga'' Pederzani & Hájek, 2005 * '' Copelatus mahleri'' Holmen & Vazirani, 1990 * '' Copelatus makokouensis'' Bilardo & Rocchi, 1995 * '' Copelatus malaisei'' Guignot, 1954 * '' Copelatus malavergnorum'' Manuel & Ramahandrison, 2022 * '' Copelatus mancus'' Sharp, 1887 * '' Copelatus marginatus'' Sharp, 1882 * ''

Copelatus marginipennis ''Copelatus marginipennis'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to ...

'' (Laporte, 1835) * ''

Copelatus martinbaehri ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hendrich, Shaverdo, Hájek & Balke, 2019 * ''

Copelatus martini ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Manuel, Deler-Hernández, Megna & Hájek, 2018 * '' Copelatus masculinus'' Régimbart, 1899 * '' Copelatus massaicus'' Guignot, 1941 * '' Copelatus mathani'' Guignot, 1952 * '' Copelatus maushomi'' Sheth, Ghate & Hájek, 2018 * '' Copelatus mbokoensis'' Bilardo & Rocchi, 2006 * '' Copelatus melanogrammus'' Régimbart, 1883 * ''

Copelatus mendax ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2011 * '' Copelatus miaowangi'' Hájek, Jiang & Jia, 2022 * ''

Copelatus minimus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 1995 * '' Copelatus minor'' Bilardo & Pederzani, 1978 * '' Copelatus minutissimus'' J. Balfour-Browne, 1939 * '' Copelatus mkambati'' Bilton & Mlambo, 2022 * '' Copelatus mocquerysi'' Régimbart, 1895 * '' Copelatus mohelicus'' Guignot, 1959 * '' Copelatus monticola'' Guignot, 1951 * '' Copelatus montivagus'' Young, 1942 * ''

Copelatus mopanshanensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Jiang, Zhao & Hájek, 2022 * '' Copelatus mulangensis'' Bameul, 2003 * '' Copelatus mundus'' Sharp, 1882 * '' Copelatus mutabilis'' Bilardo & Rocchi, 1999 * '' Copelatus mvoungensis'' Bilardo & Rocchi, 2004 * '' Copelatus mysorensis'' Vazirani, 1970 * '' Copelatus nakamurai'' Guéorguiev, 1970 * '' Copelatus nangaensis'' Guignot, 1952 * ''

Copelatus nanlingensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Jiang & Jia, 2022 * '' Copelatus nauclerus'' Guignot, 1939 * '' Copelatus neavei'' J. Balfour-Browne, 1950 * '' Copelatus neelumae'' Vazirani, 1973 * ''

Copelatus neogestroi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Balke, 2008 * '' Copelatus neoguineensis'' Zimmermann, 1919 * ''

Copelatus nguemai ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2011 * '' Copelatus nigricans'' Sharp, 1882 * '' Copelatus nigrolineatus'' Sharp, 1882 * '' Copelatus nigropennis'' Zimmermann, 1927 * '' Copelatus nigrostriatus'' Régimbart, 1895 * '' Copelatus nilssoni'' Wewalka, 1982 * '' Copelatus nimbaensis'' Legros, 1958 * '' Copelatus nitens'' Bilardo & Rocchi, 1999 * '' Copelatus nitidus'' Sharp, 1882 * '' Copelatus nodieri'' Régimbart, 1895 * '' Copelatus normalis'' Erichson, 1847 * '' Copelatus notius'' Omer-Cooper, 1965 * '' Copelatus nzei'' Bilardo & Rocchi, 1999 * '' Copelatus oblitus'' Sharp, 1882 * '' Copelatus obscurus'' Sharp, 1882 * '' Copelatus occultus'' Bilardo & Rocchi, 1995 * '' Copelatus ogasawarensis'' Kamiya, 1932 * '' Copelatus onorei'' Pederzani & Rocchi, 1982 * '' Copelatus ornatipennis'' Zimmermann, 1926 * '' Copelatus owas'' Régimbart, 1895 * '' Copelatus pachys'' Guignot, 1955 * '' Copelatus pallidus'' Régimbart, 1895 * '' Copelatus paludorum'' Guignot, 1952 * '' Copelatus pandanorum'' Scott, 1912 * '' Copelatus pantosi'' Guignot, 1958 * '' Copelatus papuensis'' J. Balfour-Browne, 1939 * '' Copelatus parabaptus'' Guignot, 1955 * '' Copelatus parallelipipedus'' Régimbart, 1895 * '' Copelatus parallelus'' Zimmermann, 1920 * ''

Copelatus pardii ''Copelatus pardii'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive", ...

'' Rocchi, 1990 * '' Copelatus parisii'' Guignot, 1959 * '' Copelatus parumstriatus'' Gschwendtner, 1934 * '' Copelatus paryphes'' Guignot, 1955 * '' Copelatus pederzanii'' Bilardo & Rocchi, 1995 * '' Copelatus pereirai'' Guignot, 1955 * '' Copelatus peridinus'' Guignot, 1955 * '' Copelatus pinnifer'' Guignot, 1952 * '' Copelatus piriensis'' Omer-Cooper, 1965 * '' Copelatus platynotus'' Régimbart, 1895 * '' Copelatus polystrigus'' Sharp, 1882 *†'' Copelatus ponomarenkoi'' Říha, 1974 * ''

Copelatus portior ''Copelatus portior'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by Félix Guignot Félix Guignot was a French physician and entomologist, born ...

'' Guignot, 1956 * '' Copelatus posticatus'' (Fabricius, 1801) * '' Copelatus poungai'' Bilardo & Rocchi, 2006 *†'' Copelatus predaveterus'' K.B. Miller, 2003 * '' Copelatus prolixus'' Sharp, 1882 * '' Copelatus prolongatus'' Sharp, 1882 * '' Copelatus propino'' Guignot, 1960 * '' Copelatus propinquus'' Régimbart, 1895 * ''

Copelatus proximus ''Copelatus proximus'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by Sharp in 1882.Copelatus pseudostriatus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * '' Copelatus pulchellus'' (Klug, 1834) * '' Copelatus pulicarius'' Régimbart, 1895 * '' Copelatus pumilus'' Régimbart, 1895 * '' Copelatus punctatus'' Bilardo & Rocchi, 1995 * '' Copelatus punctulatus'' Aubé, 1838 NA * ''

Copelatus puzhelongi ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Jiang, Hájek & Jia, 2022 * '' Copelatus quadrisignatus'' Régimbart, 1877 * ''

Copelatus quadristriatus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Pederzani & Schizzerotto, 2017 * '' Copelatus racenisi'' Guignot, 1951 * '' Copelatus ragazzii'' Régimbart, 1887 * '' Copelatus regimbarti'' Branden, 1884 * '' Copelatus restrictus'' Sharp, 1882 * '' Copelatus rimosus'' Guignot, 1952 * '' Copelatus rivalis'' Guignot, 1952 * '' Copelatus rocchii'' Bilardo, 1982 * '' Copelatus romani'' Zimmermann, 1924 * ''

Copelatus rosulae ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Jiang & Jia, 2022 * '' Copelatus royi'' Legros, 1958 * '' Copelatus rubiginosus'' Guignot, 1954 * '' Copelatus ruficapillus'' Régimbart, 1895 * '' Copelatus ruteri'' Legros, 1954 * '' Copelatus saegeri'' Guignot, 1955 * ''

Copelatus safiotra ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * '' Copelatus sahlbergi'' J. Balfour-Browne, 1939 * '' Copelatus sallaei'' Sharp, 1882 * '' Copelatus sanfilippoi'' Bilardo & Pederzani, 1972 * ''

Copelatus saverii ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Pederzani & Schizzerotto, 2017 * '' Copelatus scalptus'' Guignot, 1954 * '' Copelatus scaphites'' Guignot, 1955 * '' Copelatus schaefferi'' Young, 1942 * '' Copelatus schereri'' Wewalka, 1981 * '' Copelatus schuhi'' Hendrich & Balke, 1998 * '' Copelatus scutatus'' Guignot, 1952 * '' Copelatus scytalotus'' Guignot, 1956 * '' Copelatus senegalensis'' Lagar & Hernando, 1991 * '' Copelatus sexstriatus'' Sharp, 1882 * '' Copelatus sharpi'' Branden, 1884 * '' Copelatus sibelaemontis'' Hájek, Hendrich, Hawlitschek & Balke, 2010 * '' Copelatus silvestrii'' Régimbart, 1903 * '' Copelatus simoni'' Régimbart, 1889 * '' Copelatus singularis'' Bilardo & Rocchi, 1995 * '' Copelatus sociennus'' J. Balfour-Browne, 1952 * '' Copelatus solitarius'' Sharp, 1882 * '' Copelatus sordidipennis'' Régimbart, 1895 * '' Copelatus speciosus'' Régimbart, 1892 * ''

Copelatus spoliatus ''Copelatus spoliatus'' is a species of diving beetle. It is part of the subfamily Copelatinae in the family Dytiscidae. It was described by Félix Guignot Félix Guignot was a French physician and entomologist, born on 16 November 1882 in Av ...

'' Guignot, 1955 *†'' Copelatus stavropolitanus'' Říha, 1974 * '' Copelatus stillicidicola'' Bilardo & Rocchi, 1995 * '' Copelatus striatellus'' Boheman, 1848 * '' Copelatus striaticollis'' Lucas, 1857 * '' Copelatus striatopterus'' Aubé, 1838 * '' Copelatus striatulus'' Aubé, 1838 * '' Copelatus strigipennis'' (Laporte, 1835) * '' Copelatus strigosulus'' (Fairmaire, 1878) * '' Copelatus strinatii'' Guignot, 1958 * '' Copelatus striolatus'' Peschet, 1917 * '' Copelatus stygis'' Guignot, 1958 * '' Copelatus subdeficiens'' Régimbart, 1902 * '' Copelatus subsimilis'' Guignot, 1958 * '' Copelatus substriatus'' Kirsch, 1873 * '' Copelatus subterraneus'' Guéorguiev, 1978 * '' Copelatus sudrei'' Bameul, 2003 * '' Copelatus sulcatus'' Sharp, 1882 * '' Copelatus sulcipennis'' (Laporte, 1835) * '' Copelatus sumbawensis'' Régimbart, 1899 * '' Copelatus suppar'' Guignot, 1956 * '' Copelatus supplementaris'' Régimbart, 1895 * ''

Copelatus sylvaticoides ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2013 * '' Copelatus sylvaticus'' Guignot, 1952 * '' Copelatus takakurai'' Satô, 1985 * '' Copelatus tanaus'' Guignot, 1953 * '' Copelatus taprobanicus'' Wewalka & Vazirani, 1985 * '' Copelatus tenebrosus'' Régimbart, 1880 * '' Copelatus tengchongensis'' Hájek, Jiang & Jia, 2022 * '' Copelatus teranishii'' Kamiya, 1938 * '' Copelatus terminalis'' Sharp, 1882 * '' Copelatus ternatensis'' Régimbart, 1899 * ''

Copelatus terzanii ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2016 * '' Copelatus thiriouxi'' Peschet, 1917 * '' Copelatus thrasys'' Guignot, 1952 * '' Copelatus tibialis'' Sharp, 1882 * '' Copelatus tinctor'' Guignot, 1954 * '' Copelatus togoensis'' Régimbart, 1895 * '' Copelatus tomokunii'' Satô, 1985 * ''

Copelatus tondangoyei ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2013 * '' Copelatus tostus'' J. Balfour-Browne, 1950 * ''

Copelatus trifilis ''Copelatus trifilis'' is a species of diving beetle. It is part of the genus ''Copelatus'' in the subfamily Copelatinae of the family Dytiscidae. It was described by Félix Guignot Félix Guignot was a French physician and entomologist, born ...

'' Guignot, 1958 * '' Copelatus triglyphus'' Guignot, 1955 * '' Copelatus trilobatus'' Régimbart, 1895 * '' Copelatus tschaga'' Bilardo & Pederzani, 1972 * '' Copelatus tucuchiensis'' J. Balfour-Browne, 1939 * '' Copelatus tulagicus'' Guignot, 1942 * '' Copelatus uludanuensis'' Hendrich & Balke, 1995 * '' Copelatus undecimstriatus'' Aubé, 1838 * ''

Copelatus urceolus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Shaverdo, Hendrich & Balke, 2021 * ''

Copelatus usagarensis ''Copelatus usagarensis'' is a species of diving beetle. It is part of the subfamily Copelatinae in the family Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive", are the predaceous div ...

'' Zimmermann, 1926 * '' Copelatus vagatus'' Guignot, 1952 * '' Copelatus vagestriatus'' Zimmermann, 1919 * '' Copelatus vagus'' Bilardo & Rocchi, 1995 * '' Copelatus validus'' Sharp, 1882 * ''

Copelatus vancli ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Hendrich & Balke, 2018 * '' Copelatus vanninii'' Bilardo & Rocchi, 1999 * '' Copelatus variegatus'' Régimbart, 1895 * ''

Copelatus variistriatus ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Hájek, Shaverdo, Hendrich & Balke, 2021 * '' Copelatus venustus'' Bilardo & Rocchi, 1995 * '' Copelatus vigintistriatus'' Fairmaire, 1869 * '' Copelatus vigintisulcatus'' Régimbart, 1895 * '' Copelatus villiersi'' Guignot, 1950 * '' Copelatus virungaensis'' Bilardo, 1982 * '' Copelatus vitraci'' Legros, 1948 * '' Copelatus vivax'' Guignot, 1953 * ''

Copelatus vokoka ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * ''

Copelatus wallacei ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' J. Balfour-Browne, 1939 * '' Copelatus waltoni'' J. Balfour-Browne, 1950 * ''

Copelatus wayanadensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Manivannan & Madani, 2011 * '' Copelatus wewalkai'' Holmen & Vazirani, 1990 * '' Copelatus weymarni'' J. Balfour-Browne, 1947 * '' Copelatus xanthocephalus'' Régimbart, 1899 * ''

Copelatus xiniudong ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Jiang, Hájek & Jia, 2022 * '' Copelatus yacumensis'' Guignot, 1957 * '' Copelatus yaguarete'' Scheers, 2018 * ''

Copelatus yombensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Bilardo & Rocchi, 2015 * '' Copelatus zadiensis'' Bilardo & Rocchi, 1995 * ''

Copelatus zanabato ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Manuel & Ramahandrison, 2022 * ''

Copelatus zanatanensis ''Copelatus'' is a large genus of small diving beetles. There are some 470 described species in the genus, found worldwide, but they are most diverse in tropical South America, Africa and South-East Asia. Copelatus are often black or brown in co ...

'' Ranarilalatiana & Bergsten, 2019 * '' Copelatus zimmermanni'' Gschwendtner, 1934

References

External links

* {{Authority control Dytiscidae genera Beetles described in 1832 Beetles of Africa Beetles of Asia Beetles of Europe Beetles of North America Beetles of Australia Beetles of New Zealand